Module 6: The Multicellular Animal Body PDF

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Summary

This document details the fundamental concepts of animal form and function, focusing on the organization and structure of multicellular organisms. It explores the relationship between animal structure and its interactions with the environment.

Full Transcript

‭MODULE 6: THE MULTICELLULAR ANIMAL BODY‬ I‭nterstitial‬ ‭Fluid:‬ ‭In‬ ‭flat‬ ‭animals‬ ‭(e.g.‬ ‭tapeworms),‬ ‭ nimal‬ ‭form‬ ‭and‬ ‭function‬ ‭are‬ ‭correlated‬ ‭at‬ ‭all‬ A...

‭MODULE 6: THE MULTICELLULAR ANIMAL BODY‬ I‭nterstitial‬ ‭Fluid:‬ ‭In‬ ‭flat‬ ‭animals‬ ‭(e.g.‬ ‭tapeworms),‬ ‭ nimal‬ ‭form‬ ‭and‬ ‭function‬ ‭are‬ ‭correlated‬ ‭at‬ ‭all‬ A ‭most‬‭cells‬‭are‬‭in‬‭direct‬‭contact‬‭with‬‭its‬‭environment‬ ‭levels of organization.‬ ‭= the interstitial fluid of other animals.‬ ‭Anatomy:‬‭It is the biological form of an organism.‬ ‭ volutionary‬ ‭adaptations‬ ‭of‬ ‭more‬ c E ‭ omplex‬ ‭organisms‬ ‭enable‬ ‭sufficient‬ ‭exchange‬ ‭with‬ ‭the‬ ‭ hysiology:‬‭It‬‭is‬‭the‬‭biological‬‭functions‬‭an‬‭organism‬ P ‭environment.‬ ‭performs.‬ I‭n‬‭vertebrates,‬‭the‬‭space‬‭between‬‭cells‬‭is‬‭filled‬‭with‬ S‭ ize‬ ‭and‬ ‭shape‬ ‭affect‬‭the‬‭way‬‭an‬‭animal‬‭interacts‬ ‭interstitial‬ ‭fluid,‬ ‭which‬ ‭allows‬ ‭for‬ ‭the‬ ‭movement‬ ‭of‬ ‭with‬ ‭its‬ ‭environment.‬‭The‬‭body‬‭plan‬‭of‬‭an‬‭animal‬‭is‬ ‭material into and out of cells.‬ ‭programmed‬ ‭by‬ ‭the‬ ‭genome,‬ ‭itself‬ ‭the‬‭product‬‭of‬ ‭millions of years of evolution.‬ ‭ IERARCHICAL ORGANIZATION OF BODY PLANS‬ H ‭Most‬ ‭animals‬ ‭are‬ ‭composed‬ ‭of‬ ‭specialized‬ ‭cells‬ ‭organized into tissues that have different functions.‬ ‭ VOLUTION OF ANIMAL SIZE AND SHAPE‬ E ‭Physical‬‭laws‬‭govern‬‭strength,‬‭diffusion,‬‭movement,‬ T‭ issues‬ ‭make‬ ‭up‬ ‭organs‬‭,‬ ‭which‬ ‭together‬ ‭make‬ ‭up‬ ‭and heat exchange.‬ ‭organ systems‬‭.‬ ‭ roperties‬ ‭of‬ ‭water‬ ‭limit‬ ‭possible‬ ‭shapes‬ ‭for‬ ‭fast‬ P S‭ ome‬ ‭organs,‬ ‭such‬ ‭as‬ ‭the‬ ‭pancreas,‬ ‭belong‬ ‭to‬ ‭swimming animals.‬ ‭more than one system.‬ ‭ s‬ ‭animals‬ ‭increase‬ ‭in‬ ‭size,‬ ‭thicker‬ ‭skeletons‬ ‭are‬ A ‭required for support.‬ ‭ onvergent‬ ‭Evolution:‬ ‭results‬ ‭in‬ ‭similar‬ ‭adaptations‬ C ‭of diverse organisms facing the same challenge.‬ ‭In‬ ‭the‬ ‭ocean,‬ ‭fast‬ ‭swimmers‬ ‭have‬ ‭a‬ ‭streamlined‬ ‭fusiform shape‬‭.‬ ‭EXCHANGE WITH THE ENVIRONMENT‬ ‭ utrients,‬ ‭Waste‬ ‭Products,‬ ‭and‬ ‭Gases:‬ ‭It‬ ‭must‬ ‭be‬ N ‭exchanged‬ ‭across‬ ‭the‬ ‭cell‬ ‭membranes‬ ‭of‬ ‭animal‬ ‭cells.‬ ‭ ate‬‭of‬‭exchange:‬‭Should‬‭be‬‭proportional‬‭to‬‭a‬‭cell's‬ R ‭EXPLORING STRUCTURE AND FUNCTION IN ANIMAL‬ ‭surface‬‭area‬‭while‬‭amount‬‭of‬‭exchange‬‭material‬‭is‬ ‭TISSUES‬ ‭proportional to a cell's volume.‬ ‭Different‬ ‭tissues‬ ‭have‬ ‭different‬ ‭structures‬ ‭that‬ ‭are‬ ‭suited to their functions.‬ S‭ ingle-celled‬ ‭organism‬ ‭living‬ ‭in‬ ‭water‬ ‭has‬‭sufficient‬ ‭surface area to carry out all necessary exchange.‬ T‭ issues‬ ‭are‬ ‭classified‬ ‭into‬ ‭four‬ ‭main‬ ‭categories:‬ ‭Epithelial, Connective, Muscle, and Nervous.‬ ‭ ulticellular‬ ‭organisms‬ ‭with‬ ‭a‬ ‭saclike‬ b M ‭ ody‬ ‭plan‬ ‭have‬ ‭body‬ ‭walls‬ ‭that‬ ‭are‬ ‭only‬ ‭two‬ ‭cells‬ ‭thick,‬ ‭ pithelial‬ ‭Tissue:‬ ‭It‬ ‭covers‬ ‭the‬ ‭outside‬ ‭of‬ ‭the‬ ‭body‬ E ‭facilitating diffusion of materials.‬ ‭and‬‭lines‬‭the‬‭organs‬‭and‬‭cavities‬‭within‬‭the‬‭body.‬‭It‬ ‭9‬ i‭s‬ ‭a‬ ‭barrier‬ ‭against‬ ‭mechanical‬ ‭injury,‬ ‭pathogen‬ F‭ ibrous‬ ‭Connective‬‭Tissue:‬‭It‬‭is‬‭found‬‭in‬‭tendons‬ ‭and fluid loss [tight junction].‬ ‭(attach‬ ‭muscles‬ ‭to‬ ‭bones)‬ ‭and‬ ‭ligaments‬ ‭(connects bones at joints).‬ ‭ hape:‬ ‭Be‬ ‭cuboidal‬ ‭(like‬ ‭dice),‬ ‭columnar‬ ‭(like‬ S ‭bricks on end), or squamous (like floor tiles).‬ ‭ one:‬ ‭It‬ ‭is‬ ‭mineralized‬ ‭and‬ ‭forms‬ ‭the‬ ‭skeleton.‬ B ‭Bone cells are called Osteocytes.‬ ‭ rrangement:‬ ‭May‬ ‭be‬ ‭simple‬‭(single‬‭cell‬‭layer),‬ A ‭stratified‬ ‭(multiple‬ ‭tiers‬ ‭of‬ ‭cells),‬ ‭or‬ ‭ dipose‬ ‭Tissue:‬ ‭It‬ ‭stores‬ ‭fat‬ ‭for‬ ‭insulation‬ ‭and‬ A ‭pseudostratified‬ ‭(a‬ ‭single‬ ‭layer‬ ‭of‬ ‭cells‬ ‭of‬ ‭fuel. Its cells are called adipocytes.‬ ‭varying length).‬ ‭ lood:‬ ‭Blood‬ ‭cells‬ ‭and‬ ‭cell‬ ‭fragments‬ ‭in‬ ‭blood‬ B ‭The Polarity of Epithelia‬ ‭plasma.‬ ‭ olarity:‬ ‭It‬ ‭means‬ ‭two‬ ‭different‬ ‭faces.‬ ‭The‬ P ‭ artilage:‬ ‭It‬ ‭is‬ ‭a‬ ‭strong‬ ‭and‬ ‭flexible‬ ‭support‬ C ‭Epithelial‬ ‭has‬ ‭two‬ ‭surfaces:‬ ‭Apical‬ ‭surface‬ ‭material. Its cells are called chondrocytes.‬ ‭(outer) and Basal surface.‬ ‭ uscle‬‭Tissue:‬‭It‬‭is‬‭responsible‬‭for‬‭nearly‬‭all‬‭types‬‭of‬ M ‭ onnective‬ ‭Tissue:‬ ‭It‬ ‭mainly‬ ‭binds‬ ‭and‬ ‭supports‬ C ‭body‬‭movement.‬‭Muscle‬‭cells‬‭consist‬‭of‬‭filaments‬‭of‬ ‭other‬ ‭tissues.‬ ‭It‬ ‭contains‬ ‭sparsely‬ ‭packed‬ ‭cells‬ ‭the‬ ‭proteins‬ ‭actin‬ ‭and‬ ‭myosin,‬ ‭which‬ ‭together‬ ‭scattered‬ ‭throughout‬ ‭an‬ ‭extracellular‬ ‭matrix.‬ ‭The‬ ‭enable muscles to contract.‬ ‭matrix‬ ‭consists‬ ‭of‬ ‭fibers‬ ‭in‬ ‭a‬ ‭liquid,‬ ‭jellylike,‬ ‭or‬ ‭solid‬ ‭foundation.‬ ‭Divided in the vertebrate body into three types:‬ ‭Three Types of Connective Tissue Fiber‬ ‭All of these are made of protein.‬ ‭ keletal‬ ‭Muscle‬ ‭(Striated‬ ‭Muscle):‬ S ‭It‬ ‭is‬ ‭responsible for voluntary movement.‬ ‭ ollagenous‬ ‭Fibers:‬ ‭It‬ ‭provides‬ ‭strength‬ ‭and‬ C ‭flexibility.‬ ‭ mooth‬ ‭Muscle:‬ ‭It‬ ‭is‬ ‭responsible‬ ‭for‬ ‭involuntary‬ S ‭body activities.‬ ‭ eticular‬ ‭Fibers:‬ ‭It‬ ‭joins‬ ‭connective‬ ‭tissue‬ ‭to‬ R ‭adjacent tissues.‬ ‭ ardiac‬ ‭Muscle:‬‭It‬‭is‬‭responsible‬‭for‬‭contraction‬ C ‭of the heart.‬ ‭ lastic‬ ‭Fibers:‬ ‭It‬ ‭stretches‬ ‭and‬ ‭snaps‬ ‭back‬ ‭to‬ E ‭their original length,‬ ‭The connective tissue also contains cells, including:‬ ‭‬ ‭Fibroblasts‬‭:‬ ‭These‬ ‭secrete‬ ‭the‬ ‭protein‬ ‭of‬ ‭extracellular fibers.‬ ‭‬ ‭Macrophages‬‭:‬ ‭These‬ ‭are‬ ‭involved‬ ‭in‬ ‭the‬ ‭immune system.‬ ‭Six Major Types of Connective Tissue‬ ‭ ervous‬ ‭Tissue:‬ ‭It‬ ‭functions‬ ‭in‬ ‭the‬ ‭receipt,‬ N ‭processing,‬ ‭and‬ ‭transmission‬ ‭of‬ ‭information.‬ ‭It‬ L‭ oose‬ ‭Connective‬ ‭Tissue:‬ ‭It‬ ‭binds‬ ‭epithelia‬ ‭to‬ ‭contains:‬ ‭underlying tissues and holds organs in place.‬ ‭‬ ‭Neurons,‬ ‭or‬ ‭nerve‬ ‭cells,‬ ‭that‬ ‭transmit‬ ‭nerve‬ ‭impulses‬ ‭‬ ‭Glial cells, or glia (support cells)‬ ‭10‬ ‭ egulator:‬ ‭It‬ ‭uses‬ ‭internal‬ ‭control‬ ‭mechanisms‬ ‭to‬ R ‭control‬ ‭internal‬ ‭change‬ ‭in‬ ‭the‬ ‭face‬ ‭of‬ ‭external‬ ‭fluctuation.‬ ‭ onformer:‬ ‭It‬‭allows‬‭internal‬‭conditions‬‭to‬‭vary‬‭with‬ C ‭certain external changes.‬ ‭COORDINATION AND CONTROL‬ I‭t‬ ‭is‬ ‭in‬ ‭response‬ ‭to‬ ‭stimuli.‬ ‭It‬ ‭depends‬ ‭on‬ ‭the‬ ‭endocrine system and the nervous system.‬ ‭ ndocrine‬ ‭System:‬ ‭It‬ ‭transmits‬ ‭chemical‬ ‭signals‬ E ‭called‬ ‭hormones‬ ‭to‬ ‭receptive‬ ‭cells‬ ‭throughout‬ ‭the‬ ‭body via blood.‬ ‭ ‬ ‭hormone‬ ‭may‬ ‭affect‬ ‭one‬ ‭or‬ ‭more‬ ‭regions‬ A ‭throughout‬‭the‬‭body‬‭[depending‬‭on‬‭cells‬‭that‬‭have‬ ‭receptors for it].‬ ‭ ormones‬ ‭are‬ ‭relatively‬ ‭slow‬ ‭acting,‬‭but‬‭can‬‭have‬ H ‭ OMEOSTASIS‬ H ‭long-lasting effects.‬ ‭Used‬ ‭to‬ ‭maintain‬ ‭a‬ ‭“steady‬ ‭state”‬ ‭or‬ ‭internal‬ ‭balance regardless of external environment‬‭.‬ ‭ ervous‬ ‭System:‬ ‭It‬ ‭transmits‬ ‭information‬ ‭between‬ N ‭specific locations via the axons.‬ I‭n‬ ‭humans,‬ ‭body‬ ‭temperature,‬ ‭blood‬ ‭pH,‬ ‭and‬ ‭glucose‬ ‭concentration‬ ‭are‬ ‭each‬ ‭maintained‬ ‭at‬ ‭a‬ T‭ he‬ ‭information‬ ‭conveyed‬ ‭depends‬ ‭on‬ ‭a‬ ‭signal’s‬ ‭constant level.‬ ‭pathway,‬ ‭not‬ ‭the‬ ‭type‬ ‭of‬ ‭signal.‬ ‭Nerve‬ ‭signal‬ ‭transmission is very fast.‬ ‭ echanisms of Homeostasis‬ M ‭It‬ ‭controls‬ ‭changes‬ ‭in‬ ‭the‬ ‭internal‬ ‭environment.‬ ‭Fluctuations‬ ‭above‬ ‭or‬ ‭below‬‭a‬‭set‬‭point‬‭serve‬‭as‬‭a‬ ‭stimulus‬‭;‬‭these‬‭are‬‭detected‬‭by‬‭a‬‭sensor‬‭and‬‭trigger‬ ‭a response‬‭.‬ ‭The r‬‭esponse returns the variable to the set point‬‭.‬ ‭Feedback Control in Homeostasis‬ ‭ egative‬ ‭Feedback:‬ ‭Homeostasis‬ ‭in‬ ‭animals‬ N ‭relies‬ ‭largely‬ ‭on‬ ‭negative‬ ‭feedback,‬ ‭which‬ ‭helps to return a variable to a normal range.‬ ‭ ositive‬ ‭Feedback:‬ ‭It‬ ‭amplifies‬ ‭a‬ ‭stimulus‬ ‭and‬ P ‭does‬ ‭not‬ ‭usually‬ ‭contribute‬ ‭to‬ ‭homeostasis‬ ‭in‬ ‭The Chemical Factor‬ ‭animals.‬ ‭ nimals‬ ‭may‬ ‭regulate‬ ‭some‬ ‭environmental‬ A ‭variables while conforming to others.‬ ‭11‬ ‭ oikilotherm:‬ ‭The‬ ‭body‬ ‭temperature‬ P ‭of‬ ‭a‬ ‭poikilotherm‬‭varies‬‭with its environment.‬ ‭ omeotherm:‬ ‭The‬ ‭body‬ ‭temperature‬ ‭of‬ ‭a‬ H ‭homeotherm is relatively‬‭constant‬‭.‬ ‭ lterations in Homeostasis‬ A T‭ he‬ ‭relationship‬ ‭between‬ ‭heat‬ ‭source‬ ‭and‬ ‭body‬ ‭Set‬‭points‬‭and‬‭normal‬‭ranges‬‭can‬‭change‬‭with‬‭age‬ ‭temperature‬‭is‬‭not‬‭fixed‬‭(that‬‭is,‬‭not‬‭all‬‭poikilotherms‬ ‭or show cyclic variation.‬ ‭are ectotherms).‬ ‭‬ ‭Ectothermic‬ ‭marine‬‭fishes‬‭and‬‭invertebrates‬ ‭ ircadian‬‭Rhythm:‬‭In‬‭animals‬‭and‬‭plants,‬‭it‬‭governs‬ C ‭inhabit‬ ‭waters‬ ‭with‬ ‭such‬ ‭stable‬ ‭physiological‬ ‭changes‬ ‭that‬ ‭occur‬‭roughly‬‭every‬‭24‬ ‭temperatures.‬ ‭hours.‬ ‭Metabolic‬ ‭activities‬ ‭undergo‬ ‭daily‬ ‭cycles‬ ‭in‬ ‭‬ ‭Bats‬ ‭drop‬ ‭from‬ ‭40°C‬ ‭to‬ ‭a‬ ‭few‬ ‭degrees‬ ‭response to the circadian clock.‬ ‭above zero when they enter hibernation.‬ ‭ alancing Heat Loss and Gain‬ B ‭Organisms‬ ‭exchange‬ ‭heat‬ ‭by‬ ‭four‬ ‭physical‬ ‭processes:‬‭radiation,‬‭evaporation,‬‭convection,‬‭and‬ ‭conduction.‬ ‭HEAT REGULATION IN MAMMALS‬ ‭ cclimatization:‬ ‭A‬ ‭process‬ ‭by‬ ‭which‬ ‭homeostasis‬ A I‭t‬ ‭often‬ ‭involves‬ ‭the‬ ‭integumentary‬ ‭system‬ ‭(skin,‬ ‭can adjust to changes in the external environment.‬ ‭hair, and nails).‬ ‭ irds‬ ‭and‬ ‭mammals‬ ‭can‬ ‭vary‬ ‭their‬ ‭insulation‬ ‭to‬ B F‭ ive‬ ‭adaptations‬ ‭help‬ ‭animals‬ ‭thermoregulate:‬ ‭acclimatize to seasonal temperature changes.‬ ‭insulation,‬ ‭circulatory‬ ‭adaptations,‬ ‭cooling‬ ‭by‬ ‭evaporative‬ ‭heat‬ ‭loss,‬ ‭behavioral‬ ‭responses,‬ ‭and‬ ‭ hen‬ ‭temperatures‬ ‭are‬ ‭subzero,‬ ‭some‬‭ectotherms‬ W ‭adjusting metabolic heat production.‬ ‭produce‬ ‭“antifreeze”‬ ‭compounds‬ ‭to‬ ‭prevent‬ ‭ice‬ ‭formation in their cells.‬ I‭nsulation:‬ ‭A‬ ‭major‬ ‭thermoregulatory‬‭adaptation‬‭in‬ ‭mammals‬‭and‬‭birds.‬‭Skin,‬‭feathers,‬‭fur,‬‭and‬‭blubber‬ ‭ omeostatic‬‭processes‬‭for‬‭thermoregulation‬‭involve‬ H ‭heat flow between an animal and its environment.‬ ‭form, function, and behavior‬ ‭ irculatory‬ ‭Adaptations:‬ ‭Many‬ ‭endotherms‬ ‭and‬ C T‭ hermoregulation:‬‭It‬‭is‬‭the‬‭process‬‭by‬‭which‬‭animals‬ ‭some‬ ‭ectotherms‬ ‭can‬ ‭alter‬ ‭the‬ ‭amount‬ ‭of‬ ‭blood‬ ‭maintain‬‭an‬‭internal‬‭temperature‬‭within‬‭a‬‭tolerable‬ ‭flowing‬ ‭between‬ ‭the‬ ‭body‬ ‭core‬ ‭and‬ ‭the‬ ‭skin‬ ‭range.‬ ‭(vasodilation and vasoconstriction).‬ ‭ ndothermic:‬ ‭Animals‬ ‭can‬ ‭generate‬ ‭heat‬ ‭by‬ E T‭ he‬ ‭arrangement‬ ‭of‬ ‭blood‬ ‭vessels‬ ‭in‬ ‭many‬ ‭marine‬ ‭metabolism‬‭; birds and mammals are endotherms.‬ ‭mammals‬ ‭and‬ ‭birds‬ ‭allows‬ ‭for‬ ‭countercurrent‬ ‭exchange‬‭.‬ ‭ ctothermic:‬ ‭Animals‬ ‭gain‬ ‭heat‬ ‭from‬ ‭external‬ E ‭‬ ‭They‬ ‭transfer‬ ‭heat‬ ‭between‬ ‭fluids‬‭flowing‬‭in‬ ‭sources‬‭;‬ ‭ectotherms‬ ‭include‬ ‭most‬ ‭invertebrates,‬ ‭opposite‬ ‭directions‬ ‭and‬ ‭thereby‬ ‭reduce‬ ‭fishes, amphibians, and nonavian reptiles.‬ ‭heat loss.‬ ‭Variation in Body Temperature‬ ‭12‬ ‭ ooling‬ ‭by‬ ‭Evaporative‬ ‭Heat‬ ‭Loss:‬ ‭Many‬ ‭types‬ ‭of‬ C ‭animals‬ ‭lose‬ ‭heat‬ ‭through‬ ‭evaporation‬ ‭of‬ ‭water‬ ‭from their skin.‬ ‭ weating‬‭or‬‭Bathing:‬‭It‬‭moistens‬‭the‬‭skin,‬‭helping‬ S ‭to cool an animal down.‬ ‭ anting:‬ ‭It‬ ‭increases‬ ‭the‬ ‭cooling‬ ‭effect‬ ‭in‬ ‭birds‬ P ‭and many mammals.‬ ‭ ehavioral‬ ‭Responses:‬ ‭Both‬ ‭endotherms‬ ‭and‬ B ‭ectotherms‬ ‭use‬ ‭behavioral‬ ‭responses‬ ‭to‬ ‭control‬ ‭body temperature.‬ S‭ ome‬ ‭terrestrial‬ ‭invertebrates‬ ‭have‬ ‭postures‬ ‭that‬ ‭ nergy Allocation and Use‬ E ‭minimize or maximize absorption of solar heat.‬ ‭Organisms‬ ‭can‬ ‭be‬ ‭classified‬ ‭by‬ ‭how‬ ‭they‬ ‭obtain‬ ‭chemical energy.‬ ‭ oneybees‬‭huddle‬‭together‬‭during‬‭cold‬‭weather‬‭to‬ H ‭retain heat.‬ ‭ utotrophs:‬‭Harness‬‭light‬‭energy‬‭to‬‭build‬‭energy-rich‬ A ‭molecules (ex: plants).‬ ‭ djusting‬ A ‭Metabolic‬ ‭Heat‬ ‭Production:‬ ‭Thermogenesis‬ ‭is‬‭the‬‭adjustment‬‭of‬‭metabolic‬‭heat‬ ‭ eterotrophs:‬ ‭Harvest‬ ‭chemical‬ ‭energy‬ ‭from‬ ‭food‬ H ‭production to maintain body temperature.‬ ‭(ex: animals).‬ T‭ hermogenesis‬ ‭is‬ ‭increased‬‭by‬‭muscle‬‭activity‬‭such‬ ‭QUANTIFYING ENERGY USE‬ ‭as moving or shivering.‬ ‭ etabolic‬ ‭Rate:‬ ‭It‬ ‭is‬ ‭the‬ ‭amount‬ ‭of‬ ‭energy‬ ‭an‬ M ‭ hysiological Thermostats‬ P ‭animal uses in a unit of time.‬ ‭Thermoregulation‬ ‭in‬ ‭mammals‬ ‭is‬ ‭controlled‬ ‭by‬ ‭a‬ ‭region of the brain called the‬‭hypothalamus‬‭.‬ ‭It can be determined by:‬ ‭‬ ‭An animal’s heat loss‬ T‭ he‬ ‭hypothalamus‬ ‭triggers‬ ‭heat‬ ‭loss‬ ‭or‬ ‭heat‬ ‭‬ ‭The‬‭amount‬‭of‬‭oxygen‬‭consumed‬‭or‬‭carbon‬ ‭generating‬ ‭mechanisms.‬ ‭Some‬ ‭ectothermic‬ ‭dioxide produced‬ ‭organisms‬ ‭seek‬ ‭warmer‬ ‭environments‬ ‭to‬ ‭increase‬ ‭‬ ‭Measuring‬ ‭energy‬ ‭content‬ ‭of‬ ‭food‬ ‭their‬ ‭body‬ ‭temperature‬ ‭in‬ ‭response‬ ‭to‬ ‭certain‬ ‭consumed‬ ‭and‬ ‭energy‬ ‭lost‬ ‭in‬ ‭waste‬ ‭infections.‬ ‭products.‬ ‭Metabolic Rate and Thermoregulation‬ ‭ asal‬ ‭Metabolic‬ ‭Rate‬ ‭(BMR):‬ ‭Metabolic‬ ‭rate‬ ‭of‬ B ‭an‬ ‭endotherm‬ ‭at‬ ‭rest‬ ‭at‬ ‭a‬ ‭“comfortable”‬ ‭temperature.‬ ‭ tandard‬ ‭Metabolic‬ ‭Rate‬‭(SMR):‬‭Metabolic‬‭rate‬ S ‭of‬ ‭an‬ ‭ectotherm‬ ‭at‬ ‭rest‬ ‭at‬ ‭a‬ ‭specific‬ ‭temperature.‬ ‭13‬ ‭INFLUENCES ON METABOLIC RATE‬ ‭ ctotherms‬ ‭have‬ ‭much‬‭lower‬‭metabolic‬‭rates‬‭than‬ E ‭endotherms of a comparable size.‬ ‭ ther‬ ‭key‬ ‭factors:‬ ‭age,‬ ‭sex,‬ ‭size,‬ ‭activity,‬ O ‭temperature, and nutrition.‬ ‭Size and Metabolic Rate‬ ‭ etabolic‬ ‭rate‬ ‭is‬ ‭proportional‬ ‭to‬‭body‬‭mass‬‭to‬‭the‬ M ‭power of three quarters (m‬‭3/4‬‭).‬ S‭ maller‬ ‭animals‬ ‭have‬ ‭higher‬ ‭metabolic‬ ‭rates‬ ‭per‬ ‭gram‬‭than larger animals.‬ ‭ igher‬ ‭metabolic‬ ‭rate‬ ‭⇒‬ ‭leads‬ ‭to‬‭a‬‭higher‬‭oxygen‬ H ‭delivery‬ ‭rate,‬ ‭breathing‬ ‭rate,‬ ‭heart‬ ‭rate,‬ ‭and‬ ‭greater (relative) blood volume.‬ ‭Activity and Metabolic Rate‬ ‭ ctivity‬ ‭greatly‬ ‭affects‬ ‭metabolic‬ A ‭rate‬ ‭for‬ ‭endotherms and ectotherms.‬ I‭n‬‭general,‬‭the‬‭maximum‬‭metabolic‬‭rate‬‭an‬‭animal‬ ‭can‬‭sustain‬‭is‬‭inversely‬‭related‬‭to‬‭the‬‭duration‬‭of‬‭the‬ ‭activity.‬ F‭ or‬ ‭most‬ ‭terrestrial‬ ‭animals,‬ ‭the‬ ‭average‬ ‭daily‬ ‭rate‬ ‭of‬ ‭energy‬ ‭consumption‬ ‭is‬ ‭2-4‬ ‭times‬ ‭BMR‬ ‭(endotherms) or SMR (ectotherms).‬ T‭ he‬‭fraction‬‭of‬‭an‬‭animal’s‬‭energy‬‭budget‬‭devoted‬ ‭to‬‭activity‬‭depends‬‭on‬‭several‬‭factors:‬‭environment,‬ ‭behavior, size, and thermoregulation.‬ ‭14‬

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