Evolution Study Note - SBU3U PDF
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Darby McMillan
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These study notes cover the unifying concept of biology, delving into topics such as belief versus theory, structure and function, and examples and questions related to evolution. The notes also discuss main ideas surrounding evolution, mutations and biological change, and the evolution of evolutionary theory from 1700 to the present day .
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arbyMcMillan D 7Oct-urmom aclock/ Evolution Stu...
arbyMcMillan D 7Oct-urmom aclock/ Evolution StudyNote-SBU3U MainTopic Subtopics Notes eunifyingconcept Th Beliefvs.Theory eory-combinationofevidencetocreateanexplanation Th ofbiology Hypothesis(Beliefasmr.dustysays)-Noevidence,createdto betested Yapyapthiswillbeatestquestion StructureandFunction Biology-studyoflivingthings,theparts: - Describingthestructure - Describingthefunction - Reflectingonhow“structure”isideallysuitedtothe “function”(Evolution:0) Examples&Questions Howdoorgansbecomesoideallysuitedtotheirfunction? - Didthesepartsalwaysexistthisway?NUHUH - Whyaresomeorgansuseless? - Whydopartsdevelopduringembryodevelopment thendisappear? - Whyareweabletomanipulateorganismstochange? - Whyarethesedifferentspecieswiththesameparts? - Functionallysodif,structurallysosimilar Mainidea Organismschangetobecomemore“ideallysuited” utationand M Phylogenetictree treethatdisplaysevolution,showingthatorganismsthatare A BiologicalChange closeraremoresimilartoeachother. - Alsoreferredto“thetreeoflife”fromDarwin Phylogeny-thestudyofevolutionaryrelationships Mutation Mutation-Thesourceofnew“variants” - Geneticdiversityofgenepool - Genepool:theamountofdifferentgenes - Canbeharmfulorbeneficial: - Harmful:somethingthatdecreasesthe reproductionandsurvivalchance.Ex: - Beneficial:somethingthatincreasestherate ofreproductionandsurvival.Ex:sicklecell carriersthathaveincreasedresistanceto malaria. Selectivebreeding-choosingspecificvariantstodevelop specifictraits.Alsocalled“Artificialselection” - Naturalselection:somevariantsthatsurvivemoreare naturallymorepresent EvolutionofEvolution 1700-1800 Buffon - Structuresthatnolongerserveafunction Linnaeus - Organismschange Darwin(grandfather) - Allspeciesmayhaveoriginatedfromonespecies - Waslessridiculedthathisgrandsonbecause hehadtoevidence 1750-1850 Lamarck - Theenvironmentdriveschangeinorganisms - Puttinganorganisminanenvironmentthat forcesittochangetobetterfittheenv. - Acquiredtraits - Hebelievedthatatraitthatanorganism madewaspassedtochildren.Ex:ifaaron gotbuiltthenhisbabieswouldbeborn built - Individualspassontheir acquired/developedtraits,causing speciestoevolve Cuvier - Hestudiedpaleontology - Fossilization - Sedimentation,mineralization,exposure - Mostlyinmarineenvironments - Alsofoundinamber(treesap),ice, lava,as - Observationsmade: - Simplefossilsareatalldepths,complexonly atshallow - Shallowfossilsresemblecurrentspecies morethandeepones - distinctlayers,withuniquefossils - Theory:catastrophism.Major eventscausedmassextinction. - Contribution:speciesevolve,fromsimpleto complex Lyell - Hestudiedgeology - Observationsmade: - Explainedgeologicalprocesses(erosion, sedimentation),andsaidtheyoccuredat constantandsimilarrates. - Theory:uniformitarianism: geologicalisaconstantprocess thatoccursatasimilarrate - Hewasthefirsttosuggestthatthe earthwasmillionsofyearsold - Contribution:massivegeologicalfeatures takemillionsofyearstoform. 1831 CharlesDarwin - Hewasanaturalist(studiednature) - InvitedonatriponHMSBeagletoSouthAmericato observenatureandcollect - Collectedthousandsofsamples - Observations: - GalapagosIslands:specimens similartosouthamerica - Capeverde:specimenssimilarto Africa - Manyweirdspeciesonislands - Spent20yearsformulatinghistheory - Leadhimtodevelopthetheoryofevolution bynaturalselection Evidence Biogeography Thestudyofgeographicdistributionofspecies - Fossils - Obs:Uniquespeciesinisolatedplaces.More - Geology isolated=moreunique - biogeography - Inf.Speciesemigrate,getstuckinlocation, thenevolve. - Obs:proximityexplainsmoreofthevariationsthan habitat.(theclosertheyarethesimilarlikelytobe similarthansimilarhabitats) - Inf.Speciesemigrate,getstuck,evolve - Longerdistancetotraveland evolve Gould Ornithologist-apersonwhostudiesbirds - ConfirmsDarwin’sfinchesaredifferentspecies - Verysimilar..Commonancestor? 1837 - Darwinessay“ Transmutationofspecies” omologousand H Whencomparingsimilarstructuresbetweenspecies,someare.. AnalogousStructures - Analogous - Similarfunction,butdifferentstructure - Ex.wingsofbirdsandbutterflies, eyeoflobsterandfish,finsof fishesandwhales - Homologous - Differentfunction,butsimilarstructure - Ex.limbsofbirds,mammals, reptiles,amphibians Hypothesis - Homologousstructuresarisefromrecentcommon ancestry(inherited) - Analogousstructuresarenotduetocommon ancestry.Insteaddutosimilarenvironment (coevolution) Mechanism - mutation Embryology Studyofembryodevelopment - Revealsmanyhomologousstructuresthatdisappear beforebirth - Closertheyarethelongertheyretain homology estigial V - Someorgans/featuresthatprovidenousefulpurpose Organs/Features - Ex.dewclaw,appendix,earmuscles, whale/snakehipbones - Hypothesis:featuresleftoverfromancestors thatmadeuseofit Artificialselection Speciescanbeselectivelybredtoenhancedesirabletraits - g.agriculture,plants,animals,petslikedogsand E cats. - Hypothesis:Theenvironmentcoulddothesame thing.(ifwecanchangethesethings,whycan’tthe environmentaswell?) - Buthestilldidn’tknowhow? - HereadMalthus:“essayontheprinciple ofpopulation” - Moreindividualsproduced thancansurvive.Thus:intense competition.Onlythosewith featuresthatgiveacompetitive advantagewillsurvive. - Hedidn’texplainthatthisis whatcausesevolution,driving changesinfeaturestobetter suitenvironments. eTheoryof Th Developmentoftheory arwin:1844-after20yearsofresearch D EvolutionbyNatural Wallace:1858-sametheoryin2days Selection - Cameupwiththesameideawithoutevidence Presentedtogetherin1858 Darwin1858:“ontheoriginofspeciesbymeansofnatural selection” BLASPHEMYorlogic? Fitness:howwellanorganism's“fits”anenvironment Observations Inferences eoryofNatural Th Selection oreoffspring M areproduced thancansurvive opulationsdo P I ndividuals notcontinueto withina growinsize/have population alimit competefor resources oodandmany F vertimethe O otherresources population arelimited changesas advantageous heritable characteristics becomemore common generationafter generation I ndividuals S omeindividuals withinall willinherit populationsvary characteristics thatgivethema betterchanceof s urvivingand reproducing anyvariations M areheritable - ajorsocietalback-lash M - Importanceofvolumeofevidence Pointsofdispute Time I fevolutionisalongprocessandtheearthisonly6000years old,how?? - Resolvedbyradiometricdatingofrocks/fossils - Radioactivedecaybyunstableisotopes - Eg.C-14 - Each“halflife”ofanspecific isotope(eg.C-14)isthetimeit takesforhalfoftheisotopeto decayinthedaughterisotope. - Eg.C-14takes5,730years todecay.Forhalfofthe isotopetobeconverted, itwilltakethatlong.For ittobe75/25,itwilltake 14,060 TransitionalForms owarealltheselittlestepscausebigchanges?Likeawhaleto H abug?? - Slowlyresolvedbyfossilevidence Sourceofnewvariations Howdothesenewvariationsform? - Mutation(harmful,neutral,beneficial) - Crossingover,recombination(allelesbeingseparated thenrecombined) - Sexualreproduction S amespeciesondifferent H owdobothspeciesexistontheearthondifferentcontinents continents atthesametimes? - Platetectonics.Theystartedatthesameplaceon Pangea,andthensplit. Complexstructures ecauseorganismsaresocomplex,it’stoocomplicatedtohave B acreator. - Thingsbegunsimpleandevolvedtobemorecomplex slowly TypesofSelection StabilizingSelection Selectionwheretheaveragephenotypeisfavoredforsurvival. - Ex.Iftheaveragebeaksizeofahummingbirdis5cm, thenbirdsclosertothatsizewillhaveahigher chanceofsurvival. DirectionalSelection S electionthatincreasesordecreasestheaverageofatraitina population. - Ex.Iftheaveragehummingbirdbeaksizeis5cm,but ashiftintheenvironmentlikeadroughtforces longerbeakstobebetteratobtainingfood,thenit woulddriveanincreaseintheaverage. DisruptiveSelection S electionthatfavorstwoormorevariationsofatraitthat differsfromtheaverage. - Ex.Iftheaveragehummingbirdbeaksizeis5cm,but 2cmand8cmthrive,thenthesewillbeselected. SexualSelection S electionthatfavorsaphenotypethatspecificallyenhancesthe matingsuccessofanindividual. - Hummingbirdswithbrighterfeathersmightattract morepartners.Overtime,thenumberofbright featheredbirdswillincrease. SexualDysmorphism ephysicaldifferencesbetweenmalesandfemalesofthe Th samespecies. - Sexualselectionisthemechanismbehindthis. - Ex.Initiallybothmaleandfemale hummingbirdshadmostlywhitefluff,and maleswithmorecolorsweremoresexually attractive.Thenoverhundredsofyears, malebirdswouldbemorecolorfulthan females. - Onlyhappensinsomespecies. Extranotes - I fapopulationhasbeeninthesameenvironmentfor alongtime,youcanassumeitsstableselection.If not,disruptive. HardyWeinberg Allele Frequencies Remember:Foreachtrait,therearetwoalleles. - Ex.Height:Tall=Tshort=t Theproportionofaspecificalleleinapopulation. - Showshowcommonanalleleis. - Ex.HowfrequentT(tall)is.Ifitis0.8,then 80%frequency. quation: E P+Q=1 P=Thefrequencyofthedominantallele Q=Thefrequencyoftherecessiveallele lleleFrequency A I f60%ofallelesareG,and40%areg,then.. examplequestions P=0.6 Q=0.4 Genotypefrequencies TheproportionofaspecificGenotypeinapopulation - Showshowcommonaphenotypeis - Ex.HowfrequentTT,Tt,ttis. Equation: P2+2PQ+Q2=1 P2=Frequencyoftheheterozygousdominantgenotype(TT) 2PQ=Frequencyofthehomozygousgenotype(Tt) Q2=Frequencyoftheheterozygousrecessivegenotype(tt) enotypefrequencies G .62 +2(o.6)(0.4)+0.42=1 O examplequestions 0.36+0.48+0.16+1 36%=GG 48%=Gg 16%=gg BIGEXAMPLE Giveninfo: - 500totalfrogs - 375darkgreen Determine: - Findgenotypeandallelefrequencies Solve: - 500-375=125 - 75%aredarkgreen - 25%arelightgreen - Q2=.25 - P+√.25 =1 - P+0.5=1 - P=0.5 - 0.52+2(0,5)(0,5)+0.52 - GG=0.25 - Gg=0.5 - gg=0.25 enotypeproportionsin G GenePool=Thecombinationofallthegenesinapopulation. Populations - Populationsizemultipliedbytwo - Becauseeachindividualhas2 - Percentageofpopulationthathaveaspecificallele - T+t=1 Genotypesinpopulation - Percentageofpopulationthathasaspecificgenotype - TT+Tt+tt=1 Theproportionofindividualsthatareheteroandhomomust equal1. - Proportionsarethesameaspercentagesby multiplyingby100. - Ex.Proportion=0.6 - Percentage=60% enotypeproportions=calculatedbydividingtheindividuals G withacertaingenotypebythetotalpopulation - Percentageofthepopulationwithaspecificgenotype - Ex.if500totalpopulation,and150haveAa,then 500/160=0.64or64% Example all=TT,Tt T Short=tt TT+Tt+tt=100% 60%+30%+10%=100% 0.6+0.3+0.1=1 ardyWeinberg H - opulationsizeislarge P assumptions - Noselection - Nomutation - Noimmigration - Randommating andomchange(small R onstantallele C Randomchange-”geneticdrift” populations) frequencies - Thereisnoselection,nomutation,nomigration, theiralleleratios&genotyperatioswillstay relativelythesameforfuturegenerations. Fixed:Thegeneforatraitisat0%,or100%. - Onlyalbinosquirrelswith3legsdied,andnowno produced,andalbinosquirrelsarefixedat0%. Mostprobableresultswilloccurwithmultipletrials. - Allelesfrequenciescanchangerandomly(notdueto naturalselection)duetoasmallpopulation. - Becauseonesingledeadalbinosquirrel coulddriveafixedgeneifthere'sonly10 squirrels. - Ifitwasalargerpopulation,allthealbino squirrelswouldhavetodietohavethesame effect. - Withasmallpopulation,asingleorganismrepresents alargerportionofthepopulation. Bottleneckeffect - largepopulationisdrasticallyreduced A - Lossindiversity(esp.Allelesthatwerealreadyrare) - Mayresultinatotallynewratioinalleles Foundereffect - S mallgroupemigratesandformsanewpopulation - Newgrouplikehasnewalleleratio - Smallpopulationmayresult,causinggeneticdrift. Speciation microevolution echangesthatoccurinapopulationfromonegenerationto Th thenext. S peciation: - Speciationistheprocessofmacroevolution: macroevolution acroevolution:Thechangesthatoccurinapopulation,where M newspeciesareformedfromtheaccumulationofmultiple microevolutions. Species YDEFINITION(STUPIDTHING):whenagroupis B reproductivelyisolated,theyareconsideredadifferentspecies becausetheyformedanothergenepool. Whatisaspecies? I ndividualsisolatedawayfromtheirgenepool,forminganew species,butuniquelycausingphysiology(chemical)and morphology(features). ^impracticaltodetect,sowehavetraditionallyused morphologyfeatures..Shortterm:DNAanalysisismore effective,especiallywhenyoucan’ttelldifferences. eproductively R Theycan’tmatewithdifferentspecies. incompatible Reproductiveisolation Allopatricspeciation Notinsamegeographical- - Obviousphysicalbarrier - Humansonearth,humansonmarscan’t reproducebcofhowfartheyare - Nowdif.species Sympatricspeciation I nthesamegeographicallocation..Mustbeanother mechanism.. echanismsof M Prezygotic arriersthatoccurbeforefertilization: B reproductiveisolation Sexcellsnevermakecontact. (sympatricspeciation) - Fertilizationdoesnotoccur,zygoteneverproduced. - Behavioral Canbeforanyreason - Structural - Ecological(habitatpreference) - biochemical - Temporal(breedingtimes) - Behavioral(matingcalls) - Mechanical(morphologicalbarrierstofertilization) - Gameticisolation(chemicalmarkersforgametes) Postzygotic arriersthatoccurafterfertilization: B Sexcellsmakecontact,butsomethinghappens.. - Zygoticmortality(zygotedies) - Hybridinviability(zygotelives,butdiestoosoonto reproduce) - Hybridinfertility(zygotelives,liveslong,butcan reproduce) PatternsofSpeciation AdaptiveRadiation henorganismsdiversityrapidlyfromanancestralspecies W intoamultitudeofnewforms. - Fillsemptyniches/habitats,andavoidscompetition withtheirownspeciesbecauseeachformisassigned aposition(niche-onlyworksifopportunitiesare openfornewbirds). DivergentEvolution henorganismslydiversifyslowslyfromancestralspeciesinto W manynewforms. iverge-becomingmore D - Avoidscompetitionwithitsownspecies,butcan different competewithotherspecies. Convergentevolution henorganismsthataredistantlyrelatedbecomemore W similarovertime. onverge-becoming C - Whenspeciesundergoasimilarnice/selection moresimilar pressureandleadstoanalogousfeatures(notfrom sameancestor,butsimilarfeaturesfromsimilar factors)) Coevolution Whentwospeciesaredependentoneachother,achangein onespecieswillchangeinthetheotherspecies. - Commoninsymbioticrelationships. Macroevolution Gradualism modelofevolutionwhichtheorizesthatevolutionisaslow A anduniformprocess. - Lotsofaccumulationofadaptation - Lotsoftimeforfossilizationoftransitionals(afossil ofanancestorsthatexhibitstraitssimilartoits descendant) Punctuatedequilibrium “Gradualism”withspedupperiodsofevolution. - Includesrapidperiodsofenvironmentalchange, usuallyhappenstoasmallisolatedpopulation,fewer fossilizationsoftransitionals. HumanEvolution Modernvsoldhumans Modern - Complexreasoning,learningtools,communication - Largebraun,finehandmotorskills,walkupright Phylogeny - Primates - Commonancestor60mya - Monkeys,apes,humans,prosimians - Largebrains,forwardeyes,flexiblehands andfeet,armsthatrotate - Mosthaveopposablethumbs - Mosthavetails - Humansareclosestrelatedwithchimpanzees98.8% - Mostrecentcommonancestor6mya - Hominids(greatapes) - Inafrica,hominidevolution - Evidence:Fossilsrecord, morphologicaldifferences,tools - Homoerectusleavesafrica1.9myaandsurviveduntil 100,000mya - Homoneanderthalensis(oongaboongas) leavesafrica 400,000mya - Homosapienleavesafrica100,000mya - Evidenceofinterbreedingwithhomo neanderthalensisandhomosapien - Extinctionofhomoneanderthalensis Aretheystillevolving? - Dependsonintensityofselectionpressure - Eg.sickle-cellanemiaandmalariaresistance (hownecessaryitistosurvivemalaria determinestheintensityofselection pressure) - Medicalcompensationsrelieveselectionpressure - Eg.treatmentsforadisease(reduceshow necessarygenesarethanhelpfightoff disease) Adaptationsfrompast: - Obesity:tasteformorerichfoods - Diabetes:modernvs.primaldiet(10,000yearsofa wheatdiet,ormillionsofprimal) - VitDdeficiency:avoidingthesun