Summary

These notes cover various invertebrate phyla, including their characteristics, body plans, and examples. The document details cell specialization, early development, body symmetry, cephalization, and body cavity trends in animal evolution. It also includes detailed anatomical information.

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Invertebrates Notes #1: Kingdom Animalia: Invertebrates Notes #2: Phylum Porifera (sponges) Notes #3: Phylum Cnidaria (cnidarians) Notes #4: Phylum Platyhelminthes (flatworms) Notes #5: Phylum Nematoda (roundworms) Notes #6: Phylum Annelida (segmented worms) Notes #7: Phylum Mollusca (...

Invertebrates Notes #1: Kingdom Animalia: Invertebrates Notes #2: Phylum Porifera (sponges) Notes #3: Phylum Cnidaria (cnidarians) Notes #4: Phylum Platyhelminthes (flatworms) Notes #5: Phylum Nematoda (roundworms) Notes #6: Phylum Annelida (segmented worms) Notes #7: Phylum Mollusca (mollusks) Notes #8: Phylum Arthropoda (arthropods) Notes #9: Phylum Echinodermata (echinoderms) Notes #10: Invertebrate Systems #1) Kingdom Animalia: Invertebrates Kingdom Animalia Characteristics of animals: eukaryotic cells; lack cell walls multicellular heterotrophic two groups: 1) Invertebrates (no backbone) 2) Vertebrates (backbone) Invertebrate Phyla There are 8 invertebrate phyla: 1) Phylum Porifera (sponges) 2) Phylum Cnidaria (cnidarians) 3) Phylum Platyhelminthes (flatworms) 4) Phylum Nematoda (roundworms) 5) Phylum Annelida (segmented worms) 6) Phylum Mollusca (mollusks) 7) Phylum Arthropoda (arthropods) 8) Phylum Echinodermata (echinoderms) Trends in Animal Evolution Important trends in animal evolution: 1) Cell Specialization 2) Early Development 3) Body Symmetry 4) Cephalization 5) Body Cavity (coelom) 1) Cell Specialization All the different cell types are derived from 3 germ layers: 1) Ectoderm – outer layer 2) Mesoderm – middle layer 3) Endoderm – inner layer 2) Early Development In protostomes the blastopore becomes the mouth (most invertebrates) In deuterostomes the blastopore becomes the anus (echinoderms and vertebrates only) Cladogram showing deuterostome evolution 3) Body Symmetry Asymmetry – no plane of symmetry Radial symmetry – multiple planes of symmetry Bilateral symmetry – one plane of symmetry 4) Cephalization Cephalization – the evolutionary trend towards a concentration of nerves and sensory organs in the anterior (head) 5) Body Cavity The coelom is a fluid-filled body cavity that contains the internal organs. The three stages in the evolution of a body cavity: 1) Acoelom → 2) Pseudocoelom → 3) Coelom Acoelom Acoelom: no body cavity three germ layers stacked; no space e.g. flatworm Pseudocoelom Pseudocoelom: “false” body cavity partially lined by mesoderm e.g. roundworm Coelom Coelom: full body cavity completely lined by mesoderm e.g. annelid #2) Phylum Porifera (sponges) Phylum Porifera (sponges) Characteristics: Sponge body plan Latin “poros” = passage germ layers: none filter feeders (use collar body cavity: absent cells to trap food particles) body symmetry: asymmetric endoskeleton made of spongin and/or spicules of cephalization: absent calcium carbonate or silica amoeboid cells build spicules and help digest food Sponges – Examples Venus’ flower basket Bath sponge Tube sponge Anatomy of a Sponge – water flow Water flow: pore (ostia) → central cavity → osculum #3) Phylum Cnidaria (cnidarians) Class Hydrozoa Class Scyphozoa Class Anthozoa Phylum Cnidaria (cnidarians) Characteristics: Cnidarian body plan Latin “cnidaria” = nettles cell layers: two mostly carnivorous body cavity: absent tentacles bear stinging body symmetry: nematocysts radial alternate between polyp and medusa body forms cephalization: gastrovascular cavity absent (GVC) hydrostatic skeleton Body Forms: Polyp vs Medusa Polyp – flower-like Medusa – bell-shaped form; sessile form; free-swimming (=attached to bottom) Stinging Cells (cnidocytes) of the Tentacles A stinging nematocyst is ejected from a cnidocyte cell when triggered It immobilizes prey Class Hydrozoa Hydra Portuguese Man-of-War Class Scyphozoa Lion’s Mane Jellyfish Moon Jelly Sea Wasp Class Anthozoa Reef Coral Sea Anemone Sea Pen Sea Fan #4) Phylum Platyhelminthes (flatworms) Class Turbellaria Class Trematoda Class Cestoda Phylum Platyhelminthes (flatworms) Characteristics: Flatworm body plan Latin “platy” = flat, germ layers: three “helminth” = worm body cavity: acoelom unsegmented body symmetry: free-living or parasitic bilateral hydrostatic skeleton cephalization: present Class Turbellaria (turbellarians) Characteristics: free-living carnivores and scavengers live in fresh, salt water, or on land move with cilia marine turbellarian GVC Planaria example: Planaria, marine turbellarians Anatomy of Planaria One-opening digestive tract: Flames cells excrete mouth → pharynx → GVC excess water Class Trematoda (flukes) Characteristics: parasitic infect blood and organs life cycle typically involves more than one host: primary host – sexual reproduction occurs intermediate/secondary host – liver fluke asexual rep. occurs examples: blood flukes (e.g. Schistosoma Life Cycle (sequence the stages) Class Cestoda (tapeworms) Characteristics: internal parasites lack digestive tract body composed of many repeating sections called proglottids (contain eggs) the scolex (head) attaches to host’s tapeworm intestines example: tapeworm Anatomy of a Tapeworm Tapeworm infection occurs by ingesting infected meat (with larval cysts) #5) Phylum Nematoda (roundworms) Phylum Nematoda (roundworms) Characteristics: Roundworm body plan Latin “nemato” = thread germ layers: three unsegmented body cavity: digestive system has two pseudocoelom openings – a mouth and body symmetry: an anus bilateral hydrostatic skeleton cephalization: examples: free-living or present parasites Nematode Digestive System The two-opening digestive system: (the first phylum to have this) mouth → pharynx→ intestine → anus Adult Ascarid Worm Ascarid infection occurs through ingesting food or water containing Ascarid eggs Ascarid Worm Lifecycle (sequence the stages) Trichinella Trichinella infection occurs from consuming raw pork Eye Worm Eye worm is transmitted through the bite of a fly #6) Phylum Annelida (segmented worms) Class Polychaeta Class Oligochaeta Class Hirudinea Phylum Annelida (segmented worms) Characteristics: Annelid body plan Latin “annellus” = little ring germ layers: three body composed of segments separated by internal body cavity: coelom partitions body symmetry: digestive system has two bilateral openings closed circulatory system* cephalization: respiration by diffusion present across skin hydrostatic skeleton Class Polychaeta (polychaetes) Characteristics: most live in salt water pair of bristly, fleshy appendages on each sandworm segment examples: sandworm, fanworm fanworm Class Oligochaeta (oligochaetes) Characteristics: lack appendages few bristles terrestrial or fresh earthworm water examples: Tubifex (sludge worm) , earthworm Tubifex Earthworm Systems e.g. Nephridia are kidney-like structures that excrete nitrogenous wastes Earthworm Digestive System mouth → pharynx → esophagus → crop → gizzard → intestine → anus Class Hirudinea (leeches) Characteristics: lack appendages carnivores or blood- sucking external parasites most live in fresh water example: medicinal leech leech #7) Phylum Mollusca (mollusks) Class Bivalvia Class Gastropoda Class Cephalopoda Phylum Mollusca (mollusks) Characteristics: Mollusk body plan Latin “molluscus” = soft germ layers: three four body parts: body cavity: coelom 1) foot body symmetry: 2) shell (endo- or exoskeleton) bilateral 3) mantle cephalization: 4) visceral mass present digestive system with two openings Four Body Parts of a Mollusk (text pg. 586) Class Bivalvia Characteristics: two-part hinged shell wedge-shaped foot typically sessile as clam oyster adults some burrow in mud or sand examples: clam, oyster, scallop, mussel scallop mussel Class Gastropoda (gastropods) Characteristics: use broad, muscular foot in movement most have spiral, chambered shells; some snail slug lack shell distinct head aquatic or terrestrial examples: snail, slug, nudibranch, limpet nudibranch limpet Class Cephalopoda (cephalopods) Characteristics: foot is divided into tentacles closed circulatory system* octopus squid examples: octopus, squid, nautilus, cuttlefish nautilus cuttlefish Squid Anatomy #8) Phylum Arthropoda (arthropods) Subphylum Trilobita Subphylum Chelicerata Subphylum Uniramia Subphylum Crustacea Phylum Arthropoda (arthropods) Characteristics: Arthropod body plan Latin “arthro” = jointed, germ layers: three “poda” = foot exoskeleton of chitin body cavity: coelom jointed appendages body symmetry: segmented body bilateral many undergo cephalization: metamorphosis during present development open circulatory system* Subphylum Trilobita (trilobites) Characteristics: all extinct (known from fossil record) each appendage divided into a gill and a walking leg trilobite Subphylum Chelicerata (chelicerates) Characteristics: mouth parts: chelicerae (1st pair) – used for biting or sucking pedipalps (2nd pair) – horseshoe crab tick used for prey capture mite two-part body – cephalothorax and abdomen lack antennae examples: horseshoe crab, tick, mite, spider, tarantula, spider tarantula scorpion scorpion Chelicerate Body (e.g. spider) Subphylum Uniramia Characteristics of uniramians: almost all terrestrial one pair of antennae mandibles examples: centipedes, millipedes, insects Centipedes (Class Chilopoda) Characteristics: long body consisting of many segments one pair of legs per segment poison claws for feeding centipede carnivorous Millipedes (Class Diplopoda) Characteristics: long body consisting of many segments two pair of legs per segment mostly herbivorous millipede Insects (Class Insecta) Characteristics: body divided into three parts – head, thorax, abdomen three pairs of legs one or two pairs of wings some undergo complete metamorphosis (e.g. Insect – Three Part Body Grasshopper Systems e.g. Malpighian tubules are kidney- like structures that excrete nitrogenous wastes Grasshopper Respiratory System Oxygen enters through tiny spiracles and into tracheal tubes where it diffuses directly to tissues. Insect Metamorphosis Complete Incomplete metamorphosis: metamorphosis: Symbiosis in Arthropods Symbiosis is a close relationship between two species Three types: 1) mutualism – both organisms benefit 2) commensalism – one organism benefits, the other neither benefits nor is harmed 3) parasitism – one organism harms the other 1) Mutualism Aphids feed the ants and the ants protect the aphids 2) Commensalism Pseudoscorpions hitch a ride by grabbing hold of flying insects 3) Parasitism Parasitic wasps inject their eggs into a host (e.g. moth larvae) Subphylum Crustacea most are aquatic; salt water two pair of antennae mouth parts called barnacle crayfish mandibles pill bug carapace covers body e.g. crab, crayfish, pill bug, water flea, water flea crab barnacle #9) Phylum Echinodermata (echinoderms) Phylum Echinodermata (echinoderms) Characteristics: Echinoderm body plan Latin “echino” = spiny, germ layers: three “dermata” = skin body cavity: coelom deuterostomes body symmetry: radial water vascular system cephalization: respiration across skin absent gills endoskeleton Sea Lilies and Feather Stars Characteristics: filter feeders feathery arms for feeding and respiration sea lily examples: sea lily, feather star feather star Sea Stars Characteristics: star-shaped carnivorous bottom dwellers sea star mouth on lower surface, anus on top surface examples: crown-of thorns, sea star, sunstar crown-of-thorns sunstar Water Vascular System of Sea stars madreporite → stone canal → ring canal → radial canal → ampulla → tube feet Brittle Stars and Basket Stars Characteristics: small body disk most have only five arms lack an anus brittle star most are filter feeders or detritus feeders examples: brittle stars and basket stars basket star Sea Urchins and Sand Dollars Characteristics: lack arms body encased in rigid, box- like covering sea urchin covered with spines most grazing herbivores or detritus feeders examples: sea urchin, sand dollar sand dollar Sea Cucumbers Characteristics: leathery, cylindrical body with feeding tentacles on one end mostly detritus or filter feeders endoskeleton greatly sea cucumber reduced #10) Invertebrate Systems Invertebrate Systems Six major invertebrate systems: 1) skeletal system 2) digestive system 3) circulatory system 4) respiratory system 5) excretory system 6) nervous system Skeletal System exoskeleton endoskeleton hydrostatic skeleton Digestive System Circulatory System Respiratory System Excretory System Nervous System

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