Veterinary Anatomy: Head and Neck PDF
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This veterinary anatomy textbook covers the head, neck, and related structures of animals. The document focuses particularly on the headgut, pharynx, larynx, and teeth and their mastication in animals like dogs and horses. The file contains many diagrams and clinical case studies.
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2.1 Headgut, Pharynx & Larynx Prehension: Get food into mouth Domestic animal: lip, teeth, tongue, head & jaw movement Mastication: Mechanical breakdown Food mix w/ saliva Deglutition: Swallowing Initial phase: voluntary Remaining phases (pass soft pellet,...
2.1 Headgut, Pharynx & Larynx Prehension: Get food into mouth Domestic animal: lip, teeth, tongue, head & jaw movement Mastication: Mechanical breakdown Food mix w/ saliva Deglutition: Swallowing Initial phase: voluntary Remaining phases (pass soft pellet, in pharynx): involuntary Horse: X vomit under normal condition → Severe disease Musculature (muscles) of prehension: Zygomaticus (jaw): retract mouth angle Levator (up) labii superioris: elevate upper lip → draw it to 1 side Buccinator (side): keep food between upper & lower molar teeth Orbicularis oris: close lips, help to gather food Oral cavity: Start of GIT Extend: lip → pharynx ○ Pharynx: naso- & oro- Structure: tongue, teeth, salivary glands Dog: 1. Vestibule 1. Buccal cavity 2. Canine tooth 2. Palatine raphe: 1 a. Philtrum single line (midline) 3. Hard palate 3. Transverse ridges 4. Soft palate 4. Soft palate 5. Tongue 5. Medial Sulcus: help 6. Sublingual caruncle drinking water, 7. Palatoglossal arch centre of tongue 8. Palatine tonsil 6. Vestibule: entrance 9. Frenulum of GIT 1. Tongue 2. Hard palate: separate nasal & oral cavity (ceiling) 3. Nasopharynx 4. Soft palate: separate nasal & oral pharynx 5. Oropharynx 6. Epiglottis: cartilage, close (flip & cover) trachea when swallowing → prevent water & food going through 7. Larynx 8. Trachea 9. Oesophagus (caudal) Blue: passage of air Green: page of food & water Crossover Air can come in through mouth Liquid (eg. water) can get out through nose Brachycephalic (short nostril): disease Mostly dogs X thermoregulation, filtering Extremely elongated soft pellet ○ Block larynx & trachea entrance ○ Difficulty in breathing, snoring BOAS surgery ○ Brachycephalic obstructive airway syndrome ○ Make cuts around nose, cut part of soft pellet ○ Expand potential for breathing ○ → Improve life quality, extend life expectancy Horse: 1. Tongue 2. Hard palate 3. Pharynx 4. Soft palate: elongated, might touch epiglottis 5. Epiglottis 6. Larynx 7. Trachea 8. Oesophagus: collapsed 9. Guttural pouch: blind end & dead space in skull, extension of nasopharynx Reduce head density Thermoregulation: cold air Inflammation Phonation: sound echoing X dog, cat, ruminant Musculature of deglutition (soft palate): Soft palate: elevated by tensor & levator veli palatini (muscles) → caudal free edge seal off nasopharynx: protection ○ Food X enter nasal cavity Tongue: compress food against soft palate → bolus, muscle contraction → transfer to laryngopharynx & oesophagus CN 9 (glossopharyngeal) & 10 (vagus) Salivary gland: Across headgut: cheek, tongue, lip, oesophagus, soft palate, pharynx Produce serous (watery) / mucous (viscous, foamy & bigger) / mixed secretion ○ Saliva: water, amylase, NaHCO3, antimicrobial agent, lingual lipase Innervation: ○ Sympathetic: vasoconstriction → decrease production ○ Parasympathetic: Facial (CN 7), glossopharyngeal (CN 9), trigeminal (CN 5) Increase flow → vasodilation Monostomatic Polystomatic Away from oral cavity → singular Multiple openings, adjacent → X connective duct duct Major Parotid: @ mouth, serous (horse & Zygomatic: next to eye, carnivores ruminant) / mixed (dog) only Mandibular: @ neck, mixed Sublingual: across mouth ventrally, Sublingual (caudal & rostral): mixed across mouth ventrally, mixed Minor Palatine Labial: mucous Lingual Buccal: mucous Hyoid apparatus: Larynx (voice box): between pharynx & trachea 5 diff. bone: ○ Unpaired: Basihyoid: connect both sides ○ Paired (both sides of neck): Stylohyoid: articulate w/ skull base @ petrous temporal Epihyoid Ceratohyoid Thyrohyoid: articulate w/ thyroid cartilage of larynx Hold larynx in place, attachment for pharynx & tongue → support Cartilage: ○ W/in larynx, provide rigidity & stability ○ Unpaired (3): Epiglottis Thyroid: Adam’s apple: bump of cartilage, front of throat, protect voice box Cricoid ○ Paired (1): arytenoid Cuneiform, corniculate, vocal & muscular processes Laryngeal muscle: Swallowing: elevate / depress larynx Move individual parts of larynx → breathing, phonation (vocalisation) → Ventricularis: ○ Vocal fold adduction ○ Glottis constriction → Cricoarytenoideus dorsalis: ○ Abduct arytenoid cartilage → open glottis for easy breathing → Thyroarytenoideus → Vocalis: ○ Control vocal cord (bird: syrinx) ○ Contract → air pass through → voice production Innervation to larynx: cranial & caudal laryngeal nerves (from vagus nerve CN 10) ○ Damage: roaring (horse), air pass through → noise Tongue: -glossal Fill oral cavity & oropharynx Striated muscle: mobile, voluntary contraction Hyoid bone: support (caudal) Grooming, lapping, prehension, deglutition, vocalisation Taste bud: taste, temp. sensation, mix food w/ saliva More muscular near root (caudal, back) > tip Vasculature: ○ Lingual artery (main) ○ Branches of external carotid, facial, ascending pharyngeal artery Musculature: Intrinsic: ○ Dorsal & ventral longitudinal muscle ○ W/in mouth Extrinsic (diff. directions): ○ Styloglossus: retract & elevate ○ Genioglossus: protrude & depress ○ Hyoglossus: retract & depress ○ Geniohyoideus: below tongue, draw hyoid & tongue forward Innervation: Movement: hypoglossal (CN 12) Sensory: ○ Temp., touch, pain sensation: rostral ⅔, lingual branch of trigeminal (CN 5) ○ Taste: caudal ⅓, glossopharyngeal (CN 9) Lyssa (dog): Thick cartilage Tongue remain in retracted position Water traps in small cup Papillae: small bumps on tongue, contain taste buds, help with chewing, swallowing & speech Mechanical Conical Caudal ⅓ Thick epithelium X horse Filiform Most numerous Rostral ⅔ Point caudally X taste bud, gland, lymphatic Smallest, thick keratin on stratified squamous epithelium Cat: prominent Gustatory (taste) Foliate Caudal ⅓ 8-12 in parallel fold, 1 either side of tongue midline Stratified squamous epithelium X ruminant Fungiform Rostral ⅔ Red dot Keratinised stratified squamous epithelium, w/ blood vessel Dog: panting → heat loss Vallate Lymphatics 2.2 Teeth, Sinuses & Mastication Shark: 3000 teeth Development: focal (most important) thickening w/in thickened oral epithelium Medial of labiogingival band Mesenchyme (under laminae) condense Dental lamina form ○ + Neural crest: travel around & stay Dental bud: expand, enlarge → enamel ○ Surround dental papilla → form deciduous tooth 乳齒 1st dentine formed on enamel ○ Odontoblast: continue to form dentine, displaced from enamel Primordium (precursor 先導) of permanent tooth: bud off dental lamina ○ Small mass of cells (Epithelium →) inner layer of enamel → ameloblast ○ Make enamel (Neural crest cell-derived →) dental papilla → odontoblast ○ Outer: columnar → make dentine: surround pulp → tooth ‘root’ Cementoblast (epithelial cell, distal): become follicle cell, secrete cementum (calcified, for support) around ‘root’ (Inductive) interaction between epithelial & mesenchymal precursors: Epithelial: initial signal Mesenchyme (neural crest cell): ○ Stable morphogenetic → determine tooth form (type / shape) ○ Unstable differentiation ○ Eg. molar tooth mesenchyme + limb ectoderm → ‘tooth-shaped’ cartilage Start @ crown, progress towards root Crown emerge (start erupting) → full length Eruption: Premature loss of temporary tooth → disorganised (non-occluding 遮擋) permanent teeth Temporary tooth Permanent tooth Crown fully formed → erupt Migrate into ‘socket’ of temporary tooth ○ Root X completely Increasing pressure formed Temporary tooth root: resorb overlying hard tissue → Space for root completion (bone, baby tooth) w/ osteoblast, remodel Epithelial covering w/ gum ○ Hydrostatic pressure: push tooth gradually ‘Wear’: remove epithelium into mouth ○ Periodontal ligament: metabolism → eruptive force: pull tooth upward ○ → Loosen → shed Replace: migrate into socket Mammal: diphyodont (2 sets of teeth: deciduous → permanent) Alligator (reptile): polyphyodont (> 2 sets) Dental lamina: maintained as replacement tooth Rat: monophyodont (X ever be lost) Constantly maintained Stem cell reservoir Enamel: Only in crown X repair Ameloblast: ○ Lost on eruption ○ ‘Crown’ characteristic of each tooth type ○ Projecting into Tomes’ Fibres Maintain contact w/ new (un-mineralised) enamel ○ Produce matrix (X remodel) Hardest biological material Mineralisation: enamelin, amelogenin Dentine: Major part Produce continually Repair: rate increase Innervated Mineralised matrix: type I collagen, dentine, specific proteins Odontoblast: recede 退 from newly formed surface ~ Bone, acellular Gingiva 牙齦 / 牙肉: Adhere to tooth: @ interface, between root & crown Alveolus & periodontal ligament: Periodontal ligament: suspend tooth in alveolar socket Alveolar bone: ○ 2 plates of cortical bone, separated by spongy bone ○ Fibre: bridge (w/ cementum) ○ Lined by compact bone ○ Alveoli: remodel → translation Pulp cavity: Contain blood vessels & nerves ○ Blood & nerve supply: superior & inferior alveolar (arteries, veins, nerves) ○ ≥ 1 branch in each major elevation of crown Cementum: Calcified & mineralised, surround dentine (root) Acellular X readily resorbed, X remodel → orthodontic work Contain type I collagen Form strong connection Tooth pushed against alveolar bone → move around Homodont: teeth similar in shape & size Heterodont: diff. Brachydont: Hypsodont (hypso-: death): Fixed: X further Continuous eruption: move tooth continuously eruption Worned & repaired (below gumline) Eg. carnivore: tiger Eg. herbivore (cat), dog Aradicular: Radicular: ‘Open’ rooted: X true root ‘Closed’ roots Continuously erupt & Continuous eruption, grow under gumline fixed growth Diet: enough wear → X → Form hollow, teeth clip teeth move rostrally w/ age ○ Malocclusion Eg. horse Eg. rabbit Tooth age: → Diet, weaning time, lifetime → rough aging, sexual maturity Carnivore: bigger mouth → change teeth Horse: less dependent on mother, start eating grass Deciduous Permanent Cat Incisor 3-4 weeks 3-5 months Canine 2-4 weeks 5-6 months Premolar 5-6 weeks 4-5 months Molar / 5-6 months Formula 2 (I33, C11, P32) 2 (I33, C11, P32, M11) Dog Incisor 4-6 weeks 3-5 moths Canine 3-5 weeks 5-7 months Premolar 5-6 weeks 4-6 months Molar / 4-7 months Formula* 2 (I33, C11, P33) 2 (I33, C11, P44, M23) Horse Incisor 1 1 week 2.5 years Incisor 2 4 weels 3.5 years Incisor 3 5-9 months 4.5 years Premolar (1-4) < 1 week 6 months - 4 years Molar 1-3 / 1-4 years Formula 2 (I33, P32) 2 (I33, P3/4 3, M33) Dentition: Heterodonty: diff. regions →distinct development Eruption time: diff. between breeds Gum recede w/ age → expose ‘cervical’ part (non-enamel coated surface) Herbivore: crown remain concealed below gumline ‘Contact surface’: adjacent teeth touch ‘Occlusal surface’: opposing (up & down) teeth touch Incisor: Canine: Upper: tricuspid, lower: bi-lobed Large, curved, laterally compressed Smaller rostrally Length: root > crown Single root Single root Top stone of arch, provide integrity Premolar: Molar: Irregular, closely spaced Broader Complex, larger caudally No. of cusps in all (dog) breeds: similar Discontinuous cutting crown Dog: Sharp & rounded: shearing & tearing Brachycephalic: ○ X have all canine cheek teeth → ‘fused’ roots ○ ‘Carnassial’: modified molars 裂齒 Upper premolar 4 & lower molar 1 Horse: Continuous & considerable wear ○ Enlarged surface ○ Higher crown ○ Delayed development of root ○ Complicated folding of enamel Incisor (33): ○ High crown ○ Folded enamel surface ○ Narrowing jaw → roots converge (shift rostrally) Canine (11): ○ Rudimentary 基礎 in diastema 牙間隙 ○ Size: root > crown Premolar (3/4 3), molar (33): ○ Wolf tooth: vestigial ○ Continuous surface ○ High wear, erupt continuously ○ Width: upper > lower Rodent & lagomorph: No. of teeth: reduced ○ X canine, large diastema X pulp cavity → insensitive (can clip) Ox: X upper canine & incisor → replaced by hard, fibrous dental pad Incisor X erupt continuously Lower canine: similar to incisor Premolar & molar: ○ Size: increase caudally ○ Similar to horse ○ Erupt continuously ○ Enamel ridge: noticeable Pig: Incisor: project rostrally Canine: continuously erupting ‘tusk’ → digging & tearing Premolar & molar: rounded cusp (crown) → crushing & grinding Unerupted premolar 4 ○ All occlusal surface: covered by thin coronal cementum ○ Extend onto subgingival Apical: ○ Thin cementum ○ Deposition → enamel resorption Elephant: Only 1 pair of teeth (molar): very elaborate 5 spare replacement teeth Enamel: highly folded → resistant to massive amount of wearing Mastication: Jaw articulation: allow jaw to move around ○ Temporomandibular joint ○ Mandibular symphysis Left & right meet Jaw muscle: ○ Open: digastricus Single muscle, 2 bellies ○ Close (3 sets): temporalis, masseter, (lateral & medial) pterygoids Buccinator: X close, only push food Small cheek → large gait Swallowing: jaw X actively shut → create partial vacuum Temporomandibular joint: Between condylar process of mandible & mandibular process of skull Single joint: include left & right Retro-articular process: caudal enlargement of fossa: mandibular condyle → prevent caudal dislocation Mandibular symphysis: Fibrous Between left & right half of mandible ○ Independent rotation: can move apart Dog & ruminant: biting bone / flesh, grass / stone → integrity (X break) Fine adjustment → precise occlusion Digastricus: Jaw-opening Origin: paracondylar of occipital bone Insertion: angle of mandible Single muscle, 2 bellies: ○ Caudal: 2nd pharyngeal arch (CN 7: facial) ○ Cranial: 1st pharyngeal arch (mandibular branch of CN 5: trigeminal) Masticatory muscles: Temporalis: Derive: 1st pharyngeal arch Origin: lateral of cranium 魂精 Innervation: mandibular Insertion: coronoid process branch of trigeminal (CN 5) Pull mandible dorsal > unilateral (rostral & caudal) Origin: upper jaw Insertion: lower jaw Masseter: Maxillary of skull & zygomatic arch Insertion: caudal of mandible Several divisions: unilateral & bilateral Protrude jaw Pterygoid: Origin: pterygopalatine of skull Insertion: medial of mandible Medial: ○ Close jaw: 1 sided contraction (move to left / right) Lateral: ○ Close & protrude jaw: 1 sided contraction W/ masseter: lateral translation of mandible Nasal & paranasal (side of nose) cavity: External nose: rostral end of nasal cavity Nasal plate, philtrum Flexible & cartilaginous Unmodified skin Extend into vestibule, covered by mucosa ○ Nasolacrimal duct: open Nasal cavity: Nasal septum: ○ Separate paired cavities ○ Caudal: bone (ossified), rostral: cartilage ○ Meet hard palate (cephalic: near head) ○ Length: horse (join entire hard palate, into pharynx) > dog Head conformation: affect size ○ Long head → long cavity Turbinate bone: create turbine, air move swiftly ○ Warm air before getting into lungs ○ Covered by mucosa Neighbouring: reduce size ○ Paranasal sinuses ○ Embedded part of upper cheek teeth (esp. horse) ○ Turbinate bone ○ Mucosa: line inner wall Conchae: ○ Folding of inner nasal wall (toward septum) ○ Separated by narrow clefts → compartmentalise 劃分 into meatus(es) Common, dorsal & ventral (large), middle (small) ○ Most caudal: ethmoid turbinate More complex (carnivore > herbivore) → sense of smell Paranasal sinus(es): Diverticula (outfolding) of nasal cavity, excavate (dig) skull bones ○ Extent of excavation: diff. between species Thicken border bones of nasal cavity → communicate ○ Connect: small openings, X contra-lateral (between left & right) → All domestic species: ○ Frontal: Through ethmoid meatus Horse: via caudal maxillary sinus ○ Maxillary: Through caudal / lateral of upper jaw Associated w/ hard palate, sphenoid bone, orbit, ventral concha Horse: connect to middle nasal meatus, extensive (w/ caudal & rostral parts) Frontal sinus → caudal maxillary sinus → middle nasal meatus → nasal cavity Horse: ‘rostral shift’ of cheek teeth w/ age Projection of maxillary sinuses Erupt, X grow → space Remodel → shrink in size 2.3 GI Histology Oesophagus wall: Tubuloalveolar gland: most dense cranially ○ Lubrication ○ Eg. salivary gland: secrete saliva Non-keratinised stratified squamous epithelium ○ Protective ○ X mucosal glands (in submucosa) 4 main layers: Mucosa Submucosa Tunica muscularis Serosa Villi: isolated islands of tissue Cut through when collecting sample Between epithelia: lamina propria (in mucosa) / crypt of Lieberkühn Lie in epithelium: enterocyte, goblet cells, lymphocytes Gland: simple columnar shape, for secretion Stomach: Gastric pit → gastric gland: secrete gastric juice Contain diff. types of cells → secrete variety of enzymes ○ Eg. HCl: activate pepsin (protein digesting enzyme) Chambers: ○ Horse stomach: non-glandular more proximal ○ Rumen & canine / equine stomach: Epithelium type: diff. Muscular layers: same (mucosa, submucosa, tunica muscularis, serosa) Glands: X in rumen & equine stomach, present in canine stomach Function: only mechanical in rumen & equine stomach, both mechanical & chemical in canine stomach Canine stomach Rumen Abomasum Horse stomach Epithelium Simple Stratified Simple Glandular: Non-glandular: type columnar squamous columnar simple stratified columnar squamous Mucosal Present X Present Present X glands Reticulum: Keratinised stratified squamous epithelium ○ Upper keratinised layer: protect against abrasion ○ Deeper layers: metabolise volatile fatty acid X gland ‘Honeycomb’ appearance: ○ Mucosal layer: short & tall fold ○ Meet rumen: merge into papillae Conical / secondary papillae: present on folds → rough surface: increase food breakdown Lamina muscularis: ○ Discrete bands of smooth muscle Tunica muscularis: ○ 2 thick layers: inner (circular) & outer (longitudinal) Omasum: Keratinised stratified squamous epithelium Texture: firm, size: vary Lamellae: divide lumen into narrow & uniform recesses X gland Papillae: small & lenticular (biconvex) ○ Some: large & conical ○ Circular tunica muscularis: extend into papillae of long laminae ○ Lamina muscularis: extend into papillae encircling tunica muscularis 3 smooth muscle layers → motile 能動的 Small intestine Colon Have villi X villi Absorption & digestion Only absorption: water & electrolyte Secrete enzyme X secrete & digest 2.4 Dentistry Anatomy: Crown: ○ Covered by enamel ○ Meet root @ cemento-enamel junction (CEJ, neck of tooth, X normally visible) Enamel: ○ 96% inorganic (hydroxyapatite) Hexagonal prisms / rods Held tgt by cementing organic matrix ○ 4% water & fibrous organic material ○ Highest substance in body ○ Cover exterior surface of crown only ○ Formed by ameloblasts w/in tooth bud before eruption ○ Very limited repair once erupted Pulp: ○ Living tissue ○ In pulp chamber & root canal ○ Well innervated & vascularised ○ Comprise connective tissue, nerves, lymph, blood vessels, collagen, undifferentiated reserve mesenchymal cells (eg. odontoblast) Odontoblast: line pulp cavity, branch into dentine tubules + Nerve endings → sensitive to temp. & pain Lay down dentine & reduce pulp cavity through time ○ Pulp: nourished via vessels entering & leaving root canal @ apical delta (occasionally: accessory canal) ○ Potential danger: Physical trauma: bruising, haemorrhage, pulpitis 牙髓病 Accidental overheating from polishing / scaling → pulp necrosis Tooth fracture → pulp exposure → pulpitis & pulp necrosis Loss of blood supply → ischaemic (reduced blood supply) necrosis Dentine: ○ Main supporting structure ○ 2nd hardest tissue in body ○ 70% mineral (eg. hydroxyapatite): acellular ○ 30%: water, collagen, mucopolysaccharide ○ Main structure: dentinal tubule Extend from external surface to pulp ○ Tubule: ~ 30000-40000 / mm2 Dentine exposed → transmit pain to pulp ○ Type: Primary: form before tooth eruption Secondary: form after eruption, develop w/ age from odontoblast w/in pulp, lay down in layers w/in pulp cavity Reparative / tertiary: thermal / chemical / bacterial / mechanical trauma to odontoblast, few tubules, darker in colour, very dense Root: ○ ≥ 1 root ○ Diverge point: (bi- / tri-)furcation angle ○ End: apex Apical area: site of nerve, blood vessel & lymphatic travelling into pulp → Human: single foramen → Cat & dog: multiple canal delta arrangement → Herbivore: remain open → Horse: close eventually Alveolar bone: ○ Root: encased in alveolar processes of jaws Comprise alveolar, trabecular, compact bone ○ Cribriform plate: densest bone, line alveolus Lamina Dura: seen as white line through radiography (uninterrupted: good periodontal health) Cementum: ○ Cover enamel free roots ○ Provide attachment point for periodontal ligament ○ Composition: Similar to woven bone 45-50% inorganic (hydroxyapatite) 50-55% organic Periodontal ligament: ○ Sharpey’s fibres (taut collagen fibre bundles) Inserted in & anchored to cementum & alveolar bone Gingival 牙齦 landmark: Gingival sulcus: Between tooth & free gingival margin Crevice 縫隙: surround tooth Renew every 4-6 days ○ Oral epithelium: 6-12 days Bathed in crevicular fluid: contain antibody, neutrophil, lymphocyte → immunity Normal depth: 0.5-1 mm (cat), 1-3 mm (dog) ○ > 4 mm in dog = danger ○ Active disease & attachment loss → deepen Junctional epithelium inflamed & oedematous: separate from root surface ○ Tissues infiltrated w/ primary & secondary cells ○ → Migration of polymorphonuclear granulocyte & fluid Junctional epithelium: Bottom of sulcus Control of periodontal disease Attach gingival tissue to tooth w/ hemidesmosomes Apical (towards root) extent: cemento-enamel junction Free gingiva: Form gingival margin (visible) Surround crown of tooth Attached gingiva: Tightly adherent to subgingival connective tissue & bone, via deep rete pegs Keratised: withstand stress of ripping & tearing food Cemento-enamel junction: Between anatomical crown & root Healthy: X visible ○ Visible: clear division between shiny enamel & slightly dull root cementum ○ Recession of attachment of tooth Mucogingival junction: Between soft, fleshy mucus membrane of oral cavity & tough, collagen rich gingiva Stationary ○ Hyperplasia / recession / attachment loss → gingiva change in height Interdental papilla: ○ Gingival peak between closely adjacent teeth Prevent impaction of food & debris → Preserve during surgery ○ Coronal aspect: col (indentation) Epithelium: X keratinised Function of teeth: Incisor ○ Small, single-rooted ○ Periodontal disease → mobile easily ○ → Cutting, scooping, picking up objects, grooming Canine ○ Front of mouth ○ Loss: Lower: weaken rostral mandible 顎, tongue fall out of mouth → excessive drying, trauma to tongue Upper (esp. cats): upper lip fall inward → lower canine occlude (close up) lateral to displaced lip → excoriation (wearing off skin) / puncture 刺穿 ○ → Holding prey, display, slashing & tearing when fighting, cradle (shelter) for tongue Premolar ○ Pinking shear (side) Tip of upper tooth: point to interdental space on lower jaw X → bite defect (eg. mandibular prognathism 倒及牙, brachygnathism: parrot mouth, under/overbite) ○ → Holding, carrying, breaking food into small pieces Molar ○ Dog: high masticatory force Passive bite force: 300-800 psi Sudden localised bite force (snap jaws shut): 30000-80000 psi ○ Affected by dental caries: 7% in mature dogs ○ → Grinding food into small pieces w/ flat occlusal (biting surface of the teeth) table Modified Triadan System: tooth no.ing Consistent across species Based on permanent dentition of pig ○ 11 teeth in each quadrant: 3 incisors, 1 canine, 4 premolars, 3 molars ○ Grand total: 44 teeth Quadrant: ○ 1st digit: denote ○ 1-4: permanent (恒齒), 5-8 deciduous (falling off, 乳齒) Tooth position & no.ing (dog): ○ 2nd & 3rd digit: denote position ○ Sequence start @ midline Other species: definite landmark ○ Central incisor: 01 (following: 02 & 03) ○ Canine: 04 ○ Premolar: 05-08 ○ Molar: 09-11 ○ Cat: reduced tooth no. → carnassial as landmark Upper: last premolars (108, 208) Lower: 1st molars (309, 409) Cat Horse Rabbit X 105, 205, 305, 405, 306, 406 105, 205: vestigial premolar 1 X 3rd incisor & canine 2.5 Mastication (DL) Clinical case study: Name Information Condition Diagnosis Flash Equine Drop food bolus from → Tooth root abscess Thoroughbred mouth Swelling gelding Normal ○ X-ray: X swelling & Male chewing discharge in sinuses 4 y/o Runny nose Centre of teeth: worn out Normal BCS ○ Feed composition: Losing weight concentrate Flora Canine Sunken face (concave) https://onlinelibrary.wiley.com/doi/ful Labrador l/10.1111/jvim.15373 Female → Cancer: trigeminal nerve 18 months old Temporalis muscle atrophy 萎縮 Aladdin Camelid Eyelid & lower lip https://www.researchgate.net/publicat Dromedary drooping ion/303664110_UNILATERAL_FAC camel Regularly vaccinated IAL_PARALYSIS_IN_A_DROMED Male ARY_CAMEL 13 y/o (May have) traumatic fight history → CN7 damage Face muscle (motor): depressor labii inferioris Unilateral & bilateral ear drooping, paralysis Other large animals: bovine