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The Digestive System Chapter 16 Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. https://www.youtube.com/watch?v=YN_n 8WEHg5U Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Learning Objectives Recognize the anatomy of...
The Digestive System Chapter 16 Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. https://www.youtube.com/watch?v=YN_n 8WEHg5U Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Learning Objectives Recognize the anatomy of the digestive system List the functions of the digestive system. List the structures of the oral cavity. Describe the epithelial and muscle layers of the gastrointestinal tract. Explain the process of peristalsis. Name the types of teeth found in carnivores Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. List the structures that make up the monogastric stomach and describe the function of each area. List the segments of the small and large intestine and describe the general functions of each segment. Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Mouth Teeth Pharynx Esophagus Stomach Small intestines Large intestines Anus Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. The Digestive System Digestive tract, gastrointestinal (GI) tract, alimentary canal, gut Tube that runs from the mouth to the anus; accessory digestive organs Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Species Variation Requirements for digestion and absorption of foodstuffs varies depending on the diet of the animal. Herbivores - plant-eating animals (cattle, sheep, goats) Carnivores - meat-eating animals (cats) Omnivores - animals that eat plant material and meat Monogastric animals - simple, single stomachs Ruminants - multiple mixing and fermentation compartments in addition to the stomach Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Digestive Tract Functions 1. 2. 3. 4. 5. Prehension (grasping) of food with the lips or teeth Mastication - mechanical grinding and breaking down of food (chewing) Chemical digestion of food Absorption of nutrients and water Elimination of wastes Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Digestive Tract Structure Listing layers from inner to outer Mucosa - lining of GI tract; epithelium and loose connective tissue Submucosa - dense connective tissue; may contain glands Muscle layer - outside the submucosa Serosa - outermost layer; thin, tough connective tissue. Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Digestive Tract Structure Mesentery Sheets of connective tissue Suspend digestive tube in the abdomen from the dorsal body wall Contains blood and lymph vessels and nerves that supply the GI tract Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Digestive Tract Musculature Skeletal muscle - voluntary control Found in the mouth, pharynx, the cranial part of the esophagus, and the external anal sphincter Allows the processes of chewing, mixing saliva with food, and initiation of swallowing Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Digestive Tract Musculature Smooth muscle Wall of the majority of the esophagus, the stomach, the small intestine, the large intestine, and the internal anal sphincter Primarily arranged in circular and longitudinal layers Contraction of circular muscle fibers narrows the tract Contraction of longitudinal fibers shortens the tract Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Peristalsis Circular muscle contractions Wavelike movement along the tract Propel digestive tract contents along the tube ahead of them Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Segmental Contractions Periodic circular muscle contractions Occur in different adjacent sites Mixes digestive tract contents and slows their movement through Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Buccal Cavity Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Buccal Cavity Lips, tongue, teeth, salivary glands, hard palate, soft palate, and oropharynx Lips may play role in prehension Salivary glands - produce saliva; usually three pairs with ducts that carry the saliva to the oral cavity 1. Parotid salivary glands - ventral to the ear canals 2. Mandibular salivary glands - ventral to the parotid glands at the caudal angle of the mandible 3. Sublingual salivary glands - medial to the shafts of Mosby theitemsmandible base and derived items © 2008just by Mosby, under Inc., an affiliate ofthe Elsevier Inc. Teeth Mastication - chewing; physically break down food into smaller pieces Upper arcade - contained in maxilla and the incisive bones Lower arcade - contained in the mandible Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Teeth Shape Carnivore teeth - more pointed on their occlusal surface; slightly curved toward back of mouth Good for holding prey, tearing, cutting, shredding Herbivore teeth have flat occlusal surfaces Good for grinding plant and grain material Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Types of Teeth Incisors Grasping teeth Most rostral teeth of upper and lower arcade Canines Tearing teeth Located at the corners of the incisors Longer than other teeth Pointed at the tip Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Types of Teeth Premolars Cutting teeth Rostral cheek teeth Sharp points and surfaces in carnivores Molars Grinding teeth Caudal cheek teeth Larger, flatter occlusal surfaces Used for grinding Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Teeth Terminology Lingual – Palatal – Labial – Buccal - Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Dental Formula Represents the typical number of each type of tooth found in the upper and lower arcade Tooth type designated I for incisor, C for canine, P for premolar, and M for molar Upper case for adult teeth Lower case for deciduous teeth Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Dental Formula Ruminants have no upper incisors or upper canine teeth Dental pad - flat thick connective-tissue structure on the maxilla opposite the lower incisors and canine teeth Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Dental Formula Tooth type followed by two numbers separated by a slash mark or expressed as a fraction of one number over the other First number - number of teeth in half of the upper arcade Second number - number of teeth in half of the lower arcade Total number determined by summing all the numbers and multiplying by 2 Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Dental Formulas for Several Domestic Species Species Dental Formula Canine - puppy Canine - adult Feline - kitten Feline - adult Equine - adult Porcine - adult Bovine - adult Total Number of Teeth i3/3 c1/1 p3/3 28 I3/3 C1/1 P4/4 M2/3 42 i3/3 c1/1 p3/2 26 I3/3 C1/1 P3/2 M1/1 30 I3/3 C1/1 P3-4/3 M3/3 40 or 42 I3/3 C1/1 P4/4 M3/3 44 I0/3 C0/1 P3/3 M3/3 32 Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Table 11-1 Structure of Teeth Pulp - center of tooth Blood and nerve supply enter at the apex of the tooth root Dentin surrounds and protects the tooth pulp Cementum - hard connective tissue Covers tooth root Helps fasten the tooth securely in its bony socket Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Structure of Teeth Enamel - covers the crown of the tooth Hardest, toughest tissue in the body Gingiva - epithelial tissue that composes the gums around the teeth Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Oral Cavity Functions 1. 2. Prehend the food Initiate mastication (mechanical digestion) 3. Initiate chemical digestion 4. Breaks food into smaller particles that increase the surface area available for exposure to the enzymes involved in chemical digestion Saliva - added to food as it is chewed; moistens, softens, and shapes food into a form that is more readily swallowed Prepare food for swallowing Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Esophagus Transports swallowed material from oropharynx to stomach Enters the stomach at an angle in cardia region Surrounded by cardiac sphincter muscle As stomach expands, folds of the stomach against esophagus closes the lower end of esophagus Reduces the risk for reflux In some species, the closure is strong enough to prevent reflux or vomiting (horse, rabbit) Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Monogastric Stomach Five different areas 1. Cardia 2. Fundus 3. Body 4. Pyloric antrum 5. Pylorus Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. The Stomach Functions storage of ingested food mechanical and chemical breakdown of food production of intrinsic factor for Vitamin B12 absorption in small intestine Food leaves stomach in semi-liquid form = chyme Mosby items and derived items ©©2008 byby Mosby, Inc., an affiliate of Elsevier Inc. Copyright 2016 Elsevier, Inc. Monogastric Stomach Cardia - opening from the esophagus Fundus - distensible blind pouch; expands as more food is swallowed Body - distensible middle section , is a large mixing chamber Fundus and body contain numerous glands Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Monogastric Stomach Pyloric antrum - acts like a pump grinds up swallowed food; regulates hydrochloric acid Glands contain G cells - secrete gastrin Pylorus - muscular sphincter; regulates the movement of chyme from the stomach into the duodenum Prevents backflow of duodenal contents into the stomach Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Gastric Motility Each area of the stomach has different motor functions. Fundus and body relax with swallowing of food Body of the stomach contracts to help mix food Pyloric antrum increases contractions in response to swallowing; stimulates mixing, grinding, and propulsive contractions that move food toward the pylorus Peristalsis also occurs in stomach and intestines Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Gastric Motility Parasympathetic stimulation causes fundus to relax and increases contractions in the antrum Parasympathetic stimulation involved in small intestinal motility, secretions, and blood flow Sympathetic stimulation can cause a decrease in motility - gastric atony Sympathetic stimulation decreases blood flow Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Small Intestines Divided into three segments: Duodenum Jejunum Ileum Ileocecal sphincter There is no clear demarkation between the three segments All three can perform the functions of the small intestines which are: Peristalsis Absorption of fluids and nutrients Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Small Intestines Suspended from the body wall by mesentery duodenum receives chyme through pyloric sphincter jejunum longest part most chemical digestion and absorption occur here ileum empties into large intestine at cecum (horse), at colon (dog and cat), or at cecum and colon (ruminant and pig) Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. The Omentum A double layer connects the peritoneum that links the stomach to abdominal wall From Evans H, de Lahunta A: Miller’s anatomy of the dog, ed 4, St Louis, 2013, Saunders Mosby items and derived items ©©2008 byby Mosby, Inc., an affiliate of Elsevier Inc. Copyright 2016 Elsevier, Inc. Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Structure of the Intestines Inner mucosa Villi Microvilli Submucosal Muscle Serosa Thickness of the layers changes and is dependent on the segment of the intestine Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Small Intestine Mucosa Mucosa - many folds and villi Each villus contains thousands of microvilli (brush border) Microvilli - digestive enzymes and carrier molecules embedded in cell membranes Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Small Intestinal Peristalsis Associated with the coordinated contractions of the longitudinal and circular muscle layers Stimulated by reflexes in response to dilation of a segment of the bowel Independent of the parasympathetic nervous system Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Small Intestine Motility Segmental contractions - mix intestinal contents; slow movement through intestines Help mix digestive enzymes with the intestinal contents Bring digested materials into contact with the surface of the intestinal tract for absorption Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. The Pancreas Endocrine function: pancreatic islets Exocrine function: groups of acini Acani functional unit of pancreas From Dyce KM, Sack WO, Wenseng CJG: Textbook of veterinary anatomy, ed 4, St Louis, 2010, Saunders. Mosby items and derived items ©©2008 byby Mosby, Inc., an affiliate of Elsevier Inc. Copyright 2016 Elsevier, Inc. Digestive System Functions of Pancreas Exocrine fx: Secretes bicarbonate into the duodenum Production of: Amylase-digestion of carbohydrates Protease Trypsin-break down protein Lipase-digestion of fat Helps neutralize acidity of contents and maintains the pH in the duodenum needed for proper enzyme function Endocrine fx: Produces insulin and glucagon Help regulate blood glucose levels Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Glandular Portions of the Pancreas Endocrine portion = pancreatic islets contain several different cell types beta cells secrete insulin alpha cells produce glucagon Exocrine portion = groups of acini enzymes are vital to digestion lipase, amylase, nuclease, protease anticipation of food causes increase of secretions Mosby items and derived items ©©2008 byby Mosby, Inc., an affiliate of Elsevier Inc. Copyright 2016 Elsevier, Inc. The Liver Largest digestive gland in the body multitude of functions – many not related to digestion Mosby items and derived items ©©2008 byby Mosby, Inc., an affiliate of Elsevier Inc. Copyright 2016 Elsevier, Inc. The Liver Proteins made by hepatocytes enter bloodstream through pores in adjacent sinusoids Bile is excreted by hepatocytes Mosby items and derived items ©©2008 byby Mosby, Inc., an affiliate of Elsevier Inc. Copyright 2016 Elsevier, Inc. Digestive System Functions of Liver Produces bile - bile acids, cholesterol, bilirubin Secreted into bile ducts, into hepatic duct, then to gallbladder for storage; bile is involved in digestion process (fat) Removes toxins & infectious agents that enter the body through the wall of the GI tract Stores or metabolizes nutrients absorbed from the GI tract Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. More Functions of the Liver Protein production nearly all the plasma proteins, including albumin and blood clotting proteins Conversion of amino acids into ketoacids energy production lipid synthesis Mosby items and derived items ©©2008 byby Mosby, Inc., an affiliate of Elsevier Inc. Copyright 2016 Elsevier, Inc. The Gallbladder Located in the abdomen underneath the liver and near the pancreas, Concentrates and stores bile until needed bile enters duodenum to help digest high fat meals Provides means for liver to excrete waste products even when animal is not eating Composition of bile bile salts phospholipids cholesterol bile pigments Mosby items and derived items ©©2008 byby Mosby, Inc., an affiliate of Elsevier Inc. Copyright 2016 Elsevier, Inc. Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Large Intestines Function: Recover fluids and electrolytes Store feces until evacuation Varies greatly between species Carnivores Simple tubular organ Cecum Ascending Transverse Descending Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Rectum Terminal portion of large intestine Nervous system control of motility and secretions is similar to that of the colon Numerous mucus-secreting glands lubricate and aid the passage of contents Sensory receptors detect stretching and stimulates the defecation response Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Anus Composed of internal and external muscular sphincters Internal sphincter is under autonomic control External sphincter that is under voluntary control As the rectum distends, stretch receptors in rectum wall cause partial relaxation of the internal sphincter Anal mucosal receptors increase the sense or need for defecation Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc. Review What are the functions of the digestive system. Describe the epithelial and muscle layers of the gastrointestinal tract. What is the process of peristalsis. List the structures of the oral cavity. List and give the locations of the salivary glands. Name the types of teeth found in carnivores and describe the structure of teeth. List the structures that make up the monogastric stomach and describe the function of each area. List the segments of the small and large intestine and describe the general functions of each segment. Mosby items and derived items © 2008 by Mosby, Inc., an affiliate of Elsevier Inc.