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Questions and Answers
Which characteristic distinguishes protists from prokaryotes?
Which characteristic distinguishes protists from prokaryotes?
What type of nutrition do mixotrophic protists exhibit?
What type of nutrition do mixotrophic protists exhibit?
What does primary endosymbiosis involve?
What does primary endosymbiosis involve?
Which of the following is NOT a method of reproduction in protists?
Which of the following is NOT a method of reproduction in protists?
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Which statement is true about the classification of protists?
Which statement is true about the classification of protists?
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Which characteristic is common among members of the Clade Excavata?
Which characteristic is common among members of the Clade Excavata?
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What does it mean for protists to exhibit functional diversity?
What does it mean for protists to exhibit functional diversity?
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Why is the previous idea of amitochondriates as the oldest lineage of eukaryotes now considered inaccurate?
Why is the previous idea of amitochondriates as the oldest lineage of eukaryotes now considered inaccurate?
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What type of mitochondria do diplomonads possess?
What type of mitochondria do diplomonads possess?
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Which of the following is a characteristic of parabasalids?
Which of the following is a characteristic of parabasalids?
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Which kind of organism is primarily associated with kinetoplastids?
Which kind of organism is primarily associated with kinetoplastids?
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What distinguishes euglenids from other groups?
What distinguishes euglenids from other groups?
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What type of nutritional mode can many euglenids perform?
What type of nutritional mode can many euglenids perform?
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What is the supposed origin of the clade Chromalveolata?
What is the supposed origin of the clade Chromalveolata?
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What do members of the clade Alveolata possess?
What do members of the clade Alveolata possess?
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Which of the following diseases is caused by a kinetoplastid?
Which of the following diseases is caused by a kinetoplastid?
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Which characteristic distinguishes heteromorphic generations in the alternation of generations?
Which characteristic distinguishes heteromorphic generations in the alternation of generations?
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What is a common ecological role of oomycetes?
What is a common ecological role of oomycetes?
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What is the primary component that forms the internal skeleton of radiolarians?
What is the primary component that forms the internal skeleton of radiolarians?
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What pigment gives red algae their characteristic color?
What pigment gives red algae their characteristic color?
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In the context of Rhizarians, what is the primary function of pseudopodia?
In the context of Rhizarians, what is the primary function of pseudopodia?
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Which group includes organisms that were formerly classified as fungi?
Which group includes organisms that were formerly classified as fungi?
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Which of the following correctly describes foraminiferans (forams)?
Which of the following correctly describes foraminiferans (forams)?
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What is the significance of Archaeplastida in terms of plant evolution?
What is the significance of Archaeplastida in terms of plant evolution?
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What is a reason why the kingdom Protista is no longer considered a legitimate taxon?
What is a reason why the kingdom Protista is no longer considered a legitimate taxon?
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Which process explains the evolution of plastids and mitochondria?
Which process explains the evolution of plastids and mitochondria?
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What distinguishes primary endosymbiosis from secondary endosymbiosis?
What distinguishes primary endosymbiosis from secondary endosymbiosis?
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Which of the following best describes the life cycle of the apicomplexan Plasmodium?
Which of the following best describes the life cycle of the apicomplexan Plasmodium?
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What is an important role of protists in aquatic environments?
What is an important role of protists in aquatic environments?
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Which of the following statements about protist reproduction is true?
Which of the following statements about protist reproduction is true?
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What type of generations do heteromorphic alternation of generations refer to?
What type of generations do heteromorphic alternation of generations refer to?
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Which organism demonstrates isomorphic generations in its life cycle?
Which organism demonstrates isomorphic generations in its life cycle?
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What is the primary use of the red alga Porphyra?
What is the primary use of the red alga Porphyra?
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Which group includes unicellular, colonial, and multicellular forms?
Which group includes unicellular, colonial, and multicellular forms?
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What distinguishes plasmodial slime molds from cellular slime molds?
What distinguishes plasmodial slime molds from cellular slime molds?
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What role do dinoflagellates play in ecological relationships?
What role do dinoflagellates play in ecological relationships?
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Which organism is known to cause amoebic dysentery in humans?
Which organism is known to cause amoebic dysentery in humans?
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What is a primary distinguishing feature of gymnamoebas?
What is a primary distinguishing feature of gymnamoebas?
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In which environments are most chlorophytes typically found?
In which environments are most chlorophytes typically found?
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What encompasses the Unikonta supergroup?
What encompasses the Unikonta supergroup?
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What does the term 'phagocytosis' refer to in the context of plasmodial slime molds?
What does the term 'phagocytosis' refer to in the context of plasmodial slime molds?
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Which of the following organisms are involved in harmful ecological relationships as parasites?
Which of the following organisms are involved in harmful ecological relationships as parasites?
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Study Notes
Overview: Living Small
- Even a low-power microscope reveals a diverse range of organisms in a drop of pond water
- Protists are mostly unicellular eukaryotes, containing organelles
- Protists were once classified as a kingdom but are now considered polyphyletic, because their evolutionary history is not rooted in a single ancestor
Protists/ Protozoans
- Protists are eukaryotes and have organelles, making them more complex than prokaryotes
- Most protists are unicellular, but some are colonial and multicellular
- Protists exhibit more structural and functional diversity than any other group of eukaryotes
- Single-celled protists are complex, performing all biological functions within each cell
Protists
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Nutrition: Protists display diverse nutritional strategies:
- Photoautotrophs: Contain chloroplasts for photosynthesis
- Heterotrophs: Obtain nutrients by absorbing organic molecules or ingesting larger food particles
- Mixotrophs: Combine photosynthesis and heterotrophic nutrition
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Reproduction: Protists reproduce both asexually and sexually:
- Asexual reproduction involves simple cell division
- Sexual reproduction involves processes of meiosis and fertilization
Endosymbiosis in Eukaryotic Evolution
- Much protist diversity originates from endosymbiosis, where a eukaryotic cell engulfs another living cell:
- Primary endosymbiosis: Occurs when a eukaryotic cell engulfs a prokaryotic cell, leading to organelles like chloroplasts and mitochondria
- Secondary endosymbiosis: A eukaryotic cell engulfs another eukaryote that has already undergone primary endosymbiosis
Five Supergroups of Eukaryotes
- The relationships among protist groups are constantly evolving
- One hypothesis proposes five supergroups for all eukaryotes, including protists
Clade Excavata
- Excavata contains protists with modified mitochondria and unique flagella
- Most members are asymmetrical and single-celled, with some exceptions like slime molds with limited multicellularity
- Some possess a feeding groove "excavated" from one side of their cell (Diplomonads and Parabasalids)
- Diplomonads and Parabasalids live in anaerobic environments, lack plastids, and have modified mitochondria
- Diplomonads possess modified mitochondria called mitosomes
- Diplomonads are often parasites, for example, Giardia intestinalis
- Parabasalids have reduced mitochondria called hydrogenosomes that generate some energy anaerobically, releasing hydrogen gas as a by-product
- Parabasalids include Trichomonas vaginalis, a parasite that causes the sexually transmitted infection Trichomoniasis
- Euglenozoa is a diverse group within Excavata:
- It includes predatory heterotrophs, photosynthetic autotrophs, and pathogenic parasites
- The main characteristic of this clade is a spiral or crystalline rod within their flagella, with unknown function
- This clade includes the orders Kinetoplastids and Euglenids
Kinetoplastids
- Kinetoplastids have a single mitochondrion with an organized mass of DNA called a kinetoplast
- They are widespread, flagellated parasites of all animal groups, plants and insects
- Free-living Kinetoplastids feed on bacteria in aquatic, marine and terrestrial environments
- Kinetoplastids include Trypanosoma, which causes sleeping sickness in humans
- Another pathogenic trypanosome causes Chagas’ disease
Euglenids
- Euglenids have one or two flagella that emerge from a pocket at one end of the cell
- Many Euglenids can be both autotrophic and heterotrophic, making them mixotrophs
Chromalveolates
- Chromalveolata is a monophyletic clade, likely originating from a secondary endosymbiosis event involving a red alga
- Contains important oceanic organisms
- Range from single-celled to complex multicellular taxa
- This clade includes the Alveolates and the Stramenopiles
Alveolates
- Members of the clade Alveolata contain membrane-bounded sacs (alveoli) located just under the plasma membrane
- The function of the alveoli is unknown
- Alveolates include the dinoflagellates, ciliates, and apicomplexans
Dinoflagellates
- Dinoflagellates are mostly unicellular
- Many are photosynthetic, including some that cause "red tides"
- They have two flagella and cell walls made of cellulose plates
- Some dinoflagellates produce toxins that can kill fish
Ciliates
- Ciliates are characterized by numerous short, hair-like structures called cilia
- They are heterotrophic and use cilia to move and feed
Apicomplexans
- Apicomplexans are parasites that infect animals
- They have a unique structure called an apical complex, which helps them penetrate host cells
- Apicomplexans cause diseases such as malaria and toxoplasmosis
Stramenopiles
- Stramenopiles include diatoms, golden algae, brown algae, and water molds They are characterized by the presence of hair-like structures (stramenopiles) on their flagella
- Diatoms are unicellular algae that have a glassy cell wall called a frustule
- Golden algae are a diverse group that includes photosynthetic, heterotrophic, and mixotrophic members
- Brown algae or Phaeophytes are multicellular photosynthetic protists with a complex life cycle
Brown Algae
- The pigment fucoxanthin masks the green of chlorophyll, giving brown algae their characteristic brown color
- Brown algae resemble plants and form underwater "forests"
- All are multicellular, and most are marine
- Brown algae includes many species commonly called "seaweeds"
Alternation of Generations
- Multicellular algae have evolved a variety of life cycles
- The most complex life cycles include an alternation of generations, the alternation of multicellular haploid and diploid forms
- Heteromorphic generations are structurally different, while isomorphic generations look similar
Oomycetes
- Oomycetes include water molds, white rusts, and downy mildews
- They were once considered fungi due to their morphological similarities
- Most oomycetes are decomposers or parasites of algae and fish
- Oomycetes have filaments called hyphae, which facilitate nutrient uptake
- Oomycetes can have a significant ecological impact, as in Phytophthora infestans causing potato late blight
Rhizarians
- DNA evidence supports Rhizaria as a monophyletic clade
- Rhizaria organisms (referred to as amoebas) have threadlike pseudopodia
- Amoebas move and feed by pseudopodia
- Rhizarians include forams and radiolarians
Forams
- Foraminiferans, or forams, are named for their porous shells, called tests
- Pseudopodia extend through the pores in the test
- Pseudopodia aid in swimming and feeding
- Foram tests form an extensive fossil record and are part of marine sediments
Radiolarians
- Marine protists called radiolarians have symmetrical internal skeletons made of silica
- The pseudopodia of radiolarians radiate from the central body
- Radiolarians use their pseudopodia to engulf microorganisms through phagocytosis
Archeaplastida
- Over a billion years ago, a heterotrophic protist acquired a cyanobacterial endosymbiont
- The photosynthetic descendants of this ancient protist evolved into red algae and green algae
- Land plants are descended from the green algae
- Archaeplastida is a supergroup used by some scientists and includes red algae, green algae, and land plants
Red Algae (Rhodophytes)
- Red algae appear reddish due to an accessory pigment called phycoerythrin, which masks the green of chlorophyll
- The color varies from greenish-red in shallow water to dark red or almost black in deep water
- Red algae are usually multicellular; the largest are seaweeds
- Red algae are the most abundant large algae in the coastal waters of the tropics
Green Algae
- Named for their grass-green chloroplasts
- Plants are descended from green algae
- The two main groups are chlorophytes and charophytes
- Most chlorophytes live in freshwater, but many exist in marine and some terrestrial environments
- Example of chlorophytes: unicellular Chlamydomonas
- Some chlorophytes live in damp soil, as symbionts in lichens, or in snow
Chlorophytes
- Chlorophytes include unicellular, colonial, and multicellular forms
- Most chlorophytes have complex life cycles with both sexual and asexual reproductive stages
Unikonts
- The supergroup Unikonta includes animals, fungi, and some protists that are closely related to fungi and animals
- This group includes two clades: the amoebozoans and the opisthokonts (animals, fungi, and related protists)
- The root of the eukaryotic tree remains controversial
- It is unclear whether unikonts separated from other eukaryotes relatively early or late
Amoebozoans
- Amoebozoans are amoebas that have lobe- or tube-shaped, rather than threadlike, pseudopodia
- They include slime molds, gymnamoebas, and entamoebas
Slime Molds
- Slime molds, or mycetozoans, were once thought to be fungi
- Molecular systematics places slime molds in the clade Amoebozoa
- Slime molds are divided into plasmodial slime molds and cellular slime molds, based on their unique life cycles
Plasmodial Slime Molds
- Many plasmodial slime molds are brightly pigmented, usually yellow or orange
- At one point in their life cycle, plasmodial slime molds form a mass called a plasmodium
- The plasmodium is not multicellular, but a single mass of cytoplasm containing many diploid nuclei
- It extends its pseudopodia through decomposing material, engulfing food by phagocytosis
Cellular Slime Molds
- Cellular slime molds form multicellular aggregates where cells are separated by their membranes
- Cells feed individually, but can aggregate to function as a unit when food becomes scarce
- Dictyostelium discoideum is a model organism for studying the evolution of multicellularity
Gymnamoebas
- Gymnamoebas are common unicellular amoebozoans present in soil, freshwater, and marine environments
- Most gymnamoebas are heterotrophic and actively seek and consume bacteria and other protists
- Some feed on dead organic matter
Entamoebas
- Entamoebas are parasites of vertebrates and some invertebrates
- Entamoeba histolytica causes amoebic dysentery in humans
- Dysentery: Infection of the intestines resulting in severe diarrhea
- Spread via contaminated drinking water, food, or eating utensils
Opisthokonts
- Opisthokonts include animals, fungi, and several groups of protists
- These protists are more closely related to animals and fungi than they are to other protists
Protists play key roles in ecological relationships
- Protists are found in diverse aquatic environments
- Two important roles of protists:
- Symbiotic Protists
- Photosynthetic Protists
Symbiotic Protists
- Many protists form symbiotic relationships with other species
- Some protist symbionts benefit their hosts:
- Dinoflagellates nourish coral polyps, contributing to reef development
- Hypermastigotes digest cellulose in the gut of termites
Photosynthetic Protists
- Many protists are important producers that obtain energy from the sun
- In aquatic environments, they are primary producers alongside prokaryotes
- One-third of the world's photosynthesis is performed by diatoms, dinoflagellates, multicellular algae, etc.
- The availability of nutrients can affect the concentration of protists
Domain Eukarya: Protists
- Mostly unicellular eukaryotes
- Protista is no longer considered a valid kingdom, as it is polyphyletic
- Some protists are more closely related to plants, fungi, or animals
- Unicellular (Chlamydomonas), colonial (Volvox), and multicellular species (green algae) exist
- Protists can reproduce asexually and sexually
Plastids and mitochondria evolved by endosymbiosis
- Primary vs secondary endosymbiosis
- Red and green algae underwent secondary endosymbiosis
Different Life Cycles
- Apicomplexan Plasmodium: Malaria-causing parasite
- Requires two hosts to complete its life cycle
- Ciliate Paramecium
- Conjugation (sexual reproduction)
- Cytokinesis produces four daughter cells
- Alternation of generations
- Alternation of multicellular haploid and diploid forms
- Heteromorphic generations: Structurally different (Brown algae)
- Isomorphic generations: Look similar (Ulva)
Importance of Protists
- Form mutualistic and parasitic relationships
- E.g., malaria-causing Apicomplexan Plasmodium
- Important producers in aquatic environments
You should now be able to:
- Explain why the kingdom Protista is no longer considered a valid taxon
- Explain the process of endosymbiosis and state what living organisms are likely relatives of mitochondria and plastids
- Distinguish between endosymbiosis and secondary endosymbiosis
- Name the five supergroups, list their key characteristics, and describe some representative taxa
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Description
Test your understanding of protists with this quiz, which covers their classification, structure, and nutritional strategies. Learn about unicellular eukaryotes and their diverse ways of life, including their roles in ecosystems. This quiz is essential for biology students exploring cellular organisms.