Geographic Ecology: Island Biogeography and Species Richness
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Questions and Answers

How does island size generally correlate with species richness, according to studies in geographic ecology?

  • Larger islands tend to support more species than smaller islands. (correct)
  • Species richness is inversely proportional to island area.
  • Island size has no predictable effect on species richness.
  • Smaller islands tend to have higher species richness due to edge effects.

What did MacArthur define geographic ecology as?

  • The application of ecological principles to urban planning.
  • The search for patterns of plant and animal life that can be mapped. (correct)
  • The analysis of genetic diversity within isolated populations.
  • The study of evolutionary relationships among species across different continents.

Which factor, besides area, most significantly affects species richness on islands?

  • The degree of isolation from other landmasses. (correct)
  • The average annual rainfall of the island.
  • The primary soil type found on the island.
  • The presence of specific keystone predators.

What broad geographic patterns are central to the study of geographic ecology?

<p>Island biogeography and latitudinal patterns in species richness. (D)</p> Signup and view all the answers

How did climate warming at the end of the Pleistocene affect habitat distribution in the American Southwest?

<p>It caused forest and alpine habitats to contract to the tops of high mountains. (B)</p> Signup and view all the answers

What was the focus of Arrhenius's early work that contributed to the field of geographic ecology?

<p>Investigating the relationship between area and species number. (D)</p> Signup and view all the answers

What does the concept of 'mountain islands' refer to in the context of habitat patches on continents?

<p>Isolated high-elevation habitats that species have retreated to due to climate change. (D)</p> Signup and view all the answers

Which of the following statements best describes the influence of historical processes on regional biodiversity?

<p>Long-term historical and regional processes can significantly influence species richness and diversity. (D)</p> Signup and view all the answers

How does the equilibrium model of island biogeography explain species diversity patterns on islands?

<p>Species diversity is a dynamic balance between immigration and extinction rates. (D)</p> Signup and view all the answers

What general trend is observed when comparing lake area and fish species richness, based on the studies mentioned?

<p>There is a positive relationship between lake area and fish species richness. (D)</p> Signup and view all the answers

Why might pteridophyte diversity on islands be less affected by isolation compared to bird diversity?

<p>Pteridophytes produce light spores which facilitate long-distance dispersal more effectively than birds. (A)</p> Signup and view all the answers

How does the concept of habitat islands apply to montane vegetation and mammal distributions?

<p>Montane areas act as isolated habitats, affecting the distribution and species richness of mammals; lower isolation correlates with less montane mammal species (D)</p> Signup and view all the answers

What is the expected impact on the immigration rate of new species as a new island gradually becomes populated?

<p>The rate of immigration decreases because most new arrivals will represent species already present. (A)</p> Signup and view all the answers

Based on the principles of island biogeography, which of the following scenarios would likely result in the lowest species richness?

<p>A small island located far from the mainland. (C)</p> Signup and view all the answers

According to the equilibrium model of island biogeography, what primarily determines the rate of immigration to an island?

<p>The distance of the island from the mainland or source of immigrants. (D)</p> Signup and view all the answers

Why does the equilibrium model predict that extinction rates increase with the number of species on an island?

<p>More species increase the potential for competitive interactions, potentially driving some to extinction. (D)</p> Signup and view all the answers

Lomolino et al.'s study of montane mammal species on mountaintops in the American Southwest revealed a strong negative relationship between isolation and species number. What does this suggest about the factors influencing montane mammal diversity?

<p>Immigration is a crucial process in maintaining species diversity, as reduced isolation facilitates higher immigration rates. (B)</p> Signup and view all the answers

According to the equilibrium model, which type of island would be expected to support the fewest number of species?

<p>A small island far from the mainland. (D)</p> Signup and view all the answers

According to MacArthur and Wilson's findings on Pacific Islands, what is the primary effect of isolation on bird diversity?

<p>Increased isolation reduces bird diversity. (A)</p> Signup and view all the answers

What does the concept of 'species turnover' on islands refer to, as predicted by the equilibrium model?

<p>The change in species composition on an island due to immigrations and extinctions, while the total number of species remains relatively constant. (D)</p> Signup and view all the answers

In the Simberloff and Wilson experiment, what was the significance of using mangrove islands as 'controls'?

<p>To establish a baseline for natural fluctuations in species numbers and composition without experimental manipulation. (A)</p> Signup and view all the answers

What was the primary observation made on the defaunated mangrove islands in the Simberloff and Wilson experiment?

<p>The islands quickly returned to species numbers similar to their original state, but the composition of species had changed. (D)</p> Signup and view all the answers

How did Rydin and Borgegård's findings contribute to the understanding of island biogeography?

<p>They showed that species richness is positively correlated with island area. (B)</p> Signup and view all the answers

Consider two islands: Island A is large and near the mainland, while Island B is small and far from the mainland. According to the principles of island biogeography, which of the following statements is most likely to be true regarding their equilibrium number of species?

<p>Island A will have a higher equilibrium number of species than Island B. (C)</p> Signup and view all the answers

What is the relationship between species diversity and habitat heterogeneity in diverse communities?

<p>Species in more diverse communities tend to subdivide the environment more finely. (B)</p> Signup and view all the answers

Which of the following factors contributes to the higher species richness observed in tropical environments compared to high latitude environments?

<p>More favorable environmental conditions in the tropics. (C)</p> Signup and view all the answers

According to one hypothesis, what primary factor limits tropical species compared to physical factors?

<p>Biological factors. (A)</p> Signup and view all the answers

If the rate of speciation increases and the rate of extinction decreases in an area, what is the likely outcome regarding the number of species in that area?

<p>The number of species will increase. (A)</p> Signup and view all the answers

How does greater land area in the tropics contribute to higher species richness, according to Terborgh and Rosenzweig?

<p>Organisms can disperse widely without experiencing significant environmental changes, reducing extinction rates. (D)</p> Signup and view all the answers

What effect do larger species ranges have on allopatric speciation rates?

<p>They increase allopatric speciation rates. (B)</p> Signup and view all the answers

In the context of latitudinal gradients in species richness, if immigration is discounted at large scales, what are the two primary processes that determine the number of species in an area?

<p>Speciation and extinction. (D)</p> Signup and view all the answers

Why might physically extreme environments restrict diversity?

<p>Extreme conditions favor a smaller subset of highly adapted species. (A)</p> Signup and view all the answers

Simberloff's experiment on mangrove islands primarily demonstrated what relationship between island area and species richness?

<p>Island area has a positive influence on species richness. (C)</p> Signup and view all the answers

What is a key difference observed in species accumulation between small/medium islands and large islands?

<p>Small and medium islands continue to accumulate species, whereas large islands tend to reach equilibrium. (B)</p> Signup and view all the answers

The latitudinal gradient in species richness refers to which pattern?

<p>Species richness generally increases from middle and high latitudes towards the equator. (C)</p> Signup and view all the answers

Which of the following best describes the 'Time Since Perturbation' hypothesis for latitudinal gradients in species richness?

<p>The tropics harbor more species because they are older and have experienced less frequent disturbances. (B)</p> Signup and view all the answers

According to the 'Productivity' hypothesis, how does high productivity contribute to high species richness?

<p>High productivity provides more energy to divide among populations, leading to larger populations and lower extinction rates. (B)</p> Signup and view all the answers

Why is it difficult to separate the effects of habitat diversity from area effects when studying island biogeography?

<p>Larger islands tend to have greater habitat diversity, making it hard to isolate the influence of each factor on species richness. (A)</p> Signup and view all the answers

If an island's area is reduced by 50% as part of an experimental manipulation, what is the most likely immediate effect on the island's species richness?

<p>A decrease in species richness due to the loss of habitat and resources. (C)</p> Signup and view all the answers

Which of the following statements accurately reflects Connell's (1978) proposition regarding disturbance and tropical diversity?

<p>Tropical diversity is maintained by disturbance. (A)</p> Signup and view all the answers

Flessa and Brown's research indicated which relationship between continental area and species richness?

<p>A positive relationship, where larger continental areas correlated with higher species richness. (A)</p> Signup and view all the answers

Which factors did Bond and Goldblatt attribute to the unusually high species richness in the Cape Floristic Region of South Africa?

<p>Continental drift, wide variety of soil types, and repeated expansion, contraction, and isolation of plant populations. (C)</p> Signup and view all the answers

Why do temperate forests in Europe have lower tree species richness compared to those in Eastern Asia and Eastern North America, despite having roughly equal areas?

<p>Mountain ranges in Europe oriented east-west acted as barriers, hindering southward retreat during glacial periods and increasing extinction rates. (C)</p> Signup and view all the answers

How did Rosenzweig's findings expand upon the understanding of the relationship between area and species richness?

<p>By confirming the relationship specifically within tropical regions. (A)</p> Signup and view all the answers

What conclusion can be drawn from the comparison of tree species richness in temperate forests of Europe, Eastern Asia, and Eastern North America?

<p>Historical and geographical factors can override the influence of area on species richness. (A)</p> Signup and view all the answers

In which region did Ralph (1985) find a negative correlation between foliage height diversity and bird species diversity?

<p>Argentina (C)</p> Signup and view all the answers

The Cape region of South Africa and the California Floristic Province are both biodiversity hotspots, but the Cape region exhibits higher species richness despite being smaller. What does this suggest about the factors influencing species diversity?

<p>Historical and regional factors can be more influential than area in determining species richness. (D)</p> Signup and view all the answers

Which of the following scenarios would most likely lead to increased species richness in a given region over long periods of time?

<p>A series of geographic isolations followed by periods of reconnection and expansion. (C)</p> Signup and view all the answers

Flashcards

Geographic Ecology

The study of ecological structure and process at large geographic scales.

Island Biogeography Key Concept

Species richness increases with area and decreases with isolation.

Arrhenius

Developed quantitative relationship between area and number of species.

Island Area and Species Richness

Island area is often the best single predictor of species richness.

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Latitudinal Gradients in Species Richness

A pattern where species richness decreases as you move away from the equator towards the poles.

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Preston

Number of species compared to area of west indies island

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Mountain Islands

Mountain tops becoming isolated habitat patches as climate warms

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Long-term Historical and Regional Processes

Species richness influenced over long term historical and regional processes

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Montane Habitat Islands

Montane vegetation islands formed as continuous forests fragmented, often associated with mountains.

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Lake Area and Species Richness

The number of species in a lake tends to increase as the lake's area increases.

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Island Isolation and Diversity

Isolation tends to decrease bird diversity on islands. Pteridophyte diversity is less affected due to better dispersal.

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Montane Isolation Effects

Increasing isolation from other mountaintops is correlated with lower mammal species numbers.

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Island Biogeography

The diversity of species on an island which is determined by the rates of immigration and extinction.

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Immigration Rate on new Islands

The rate of new species arriving on an island is highest when the island is new and empty.

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Declining Immigration Rate

As more species populate the island, the immigration rate declines because fewer new arrivals will be unique species.

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Equilibrium Model of Island Biogeography

Model explaining island species diversity as a balance between immigration and extinction rates.

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Equilibrium Model

A model where the number of species on an island is balanced by immigration and extinction rates.

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Extinction Rate Factors

As species number rises extinction rate increases due to smaller population sizes and more interspecific competition

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Species Number Prediction

The number of species is predicted where immigration and extinction rates intersect.

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Island Biogeography Drivers

Distance and size are the primary drivers of immigration and extinction respectively.

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Species Turnover

Species composition changes over time, despite a stable number of species.

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Island type Vs number of species

Islands closer to the mainland support higher number of species more than small far islands which support the lowest

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Stable Equilibrium Effect

Species number remains constant whiles species composition changes due to equal immigrations and extinctions

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Area and Species richness

Species richness is positively correlated with island area based on variation

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Island Area Effect

Increasing island area often leads to increased species richness.

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Simberloff's Experiment

A study where Simberloff reduced the area of mangrove islands to observe the effect on species richness.

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Latitudinal Gradient

Generally, species richness increases closer to the equator and decreases towards the poles.

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Time Since Perturbation

The idea that the tropics have had more time for speciation due to their older age and less frequent disturbances.

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Productivity Hypothesis

The hypothesis that high productivity supports high species richness by providing more energy and larger populations.

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Glaciation Impact

Glaciation disrupted middle and high latitudes, influencing species richness.

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Connell's Disturbance Hypothesis

Tropical diversity is maintained by disturbance.

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Species Richness

The number of different species living in a specific region.

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Environmental Heterogeneity Hypothesis

The idea that the tropics have more species because they contain a wider variety of environments.

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Favorableness Hypothesis

The concept that tropical environments are more stable and favorable for species survival and diversification compared to higher latitude environments.

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Biological Interactions Hypothesis

Biological factors, such as predation and parasitism, play a more significant role in limiting species in the tropics than physical factors.

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Speciation and Extinction Rates

The number of species in a region is determined by the rate at which new species are formed (speciation) minus the rate at which species disappear (extinction).

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Tropical Speciation/Extinction

Tropics have more species because they have higher rates of speciation and/or lower rates of extinction.

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Area Hypothesis

Greater tropical species richness is explained by the larger area of tropical regions.

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Species Ranges and Speciation

Describes species ranges increasing allopatric speciation rates.

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Allopatric Speciation

A process where new species arise due to geographic isolation.

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Area and Richness

Positive relationship between continental area and species richness.

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Cape Region Anomaly

The Cape region of South Africa is smaller than the California Floristic Province, yet has more species.

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Cape's Richness Factors

Continental drift, varied soil types, and population dynamics contributed to high species richness.

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Temperate Forest Diversity

Temperate forests in Europe, Eastern Asia, and Eastern North America have different diversity levels despite similar areas.

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European Mountains' Impact

East-west mountain barriers in Europe cut off southward retreat during the last ice age.

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Extinction and Richness

Higher extinction rates lead to lower species richness.

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Foliage Diversity vs. Birds

Negative correlation between foliage height diversity and bird species diversity in Argentina exists.

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Historical & Geographic Influences

Historical and geographic factors, such as continental drift & soil diversity, influence species richness.

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Study Notes

  • Geographic ecology is defined as the search for patterns of plant and animal life that can be mapped.
  • Geographic ecology studies ecological structure and process at large geographic scales.

Area, Isolation, and Species Richness

  • Arrhenius developed the quantitative relationship between area and the number of species.
  • Area and island size are related to the number of species found on the island.
  • Fewest bird species live on the smallest islands, and the most species on the largest islands.
  • Island area is the best single species richness predictor for woody plants, carabid beetles, and land snails.
  • Forest and alpine habitats contracted to mountaintops across the American Southwest due to climate warming at the end of the Pleistocene epoch.
  • Tonn and Magnuson studied Wisconsin lakes.
  • The number of species increases with the area of the lake
  • Lakes can be considered habitat islands.
  • Barbour and Brown used 70 lakes from around the world to find a positive relationship between area and fish species richness.
  • MacArthur and Wilson found that isolation reduces bird diversity on Pacific Islands.
  • Birds and pteridophytes have increased diversity with increasing island area in the Azore and Channel Islands.
  • Bird diversity is influenced by isolation, but pteridophyte diversity is not, reflecting differences in dispersal abilities (pteridophytes produce light spores).
  • Lomolino et al. looked at montane mammal species on mountaintops across the American Southwest.
  • A strong negative relationship was discovered between isolation and species number, suggesting that immigration is important in maintaining diversity.

Equilibrium Model of Island Biogeography

  • MacArthur and Wilson explained the equilibrium model of island biogeography, explaining patterns of species diversity on islands due to the balance of immigration and extinction rates.
  • Immigration rates would be highest on a new island with no organisms.
  • As species accumulate, the rate of immigration declines as fewer arrivals are new species.
  • The extinction rate rises with an increasing number of species on an island for three reasons.
  • More species increases the pool of potential extinctions.
  • As the number of species increases, the population size of each must diminish.
  • As the number of species increases, the potential for competitive interactions between species increases.
  • The point where the immigration and extinction lines cross predicts the number of species that will occur on an island.
  • Immigration rate mainly determined by source distance from source of immigrants.
  • Extinction rate is determined mainly by island size.
  • Large near islands support the highest number of species.
  • Small far islands support the lowest number of species.
  • Small near and large far islands support intermediate species numbers.
  • The equilibrium model predicts that species composition on islands is dynamic, referred to as species turnover.
  • Diamond found birds in nine Calif. Channel Islands in a stable equilibrium due to approximately equal numbers of immigrations and extinctions.
  • Species turnover occurs at a stable equilibrium.
  • Simberloff and Wilson studied insect recolonization in the Florida Keys by using two mangrove islands as controls and six mangrove islands that were defaunated.
  • The mangroves were tracked for one year.
  • The mangrove island control species number was constant and its composition changed and the defaunated islands nearly returned to original species numbers.
  • Rydin and BorgegÃ¥rd found that variation in species richness correlated positively with island area explaining 44 to 85% of species richness variation among islands.
  • Small and medium islands continued to accumulate species, whereas large islands attained equilibrium.
  • Simberloff tested the effect of island area on species richness in mangrove islands by reducing the area of some islands and leaving others as controls.
  • In cases where area was reduced, species richness decreased (and richness on the control island increased slightly).
  • Islands with reduced area lost species with each area reduction.
  • Area has a positive influence on species richness.

Latitudinal Gradients in Species Richness

  • Species richness generally increases from middle and high latitudes to the equator.
  • The pattern of increased species in the tropics is pervasive and dramatic.
  • Brown (1988) grouped hypotheses into six categories.
  • The tropics are older and disturbed less frequently, allowing for more time for speciation and reducing the extinction rate.
  • Less frequent disturbance reduces the extinction rate.
  • High productivity contributes to high species richness due to more energy to divide among populations.
  • Tropics have more species because they are more heterogeneous.
  • Tropics provide more favorable environments due to less temperature variation when compared with high latitudes.
  • Tropical species are limited more by biological factors than physical factors.
  • Number of species in the tropics reflects higher rates of speciation.
  • Terborgh and Rosenzweig proposed that a greater area would lead to more species diversity.
  • More land and water and more uniform temperatures in the tropics means organisms can disperse widely without significant changes, should reducing extinction rates, and larger species ranges should increase allopatric species rates.

Historical and Regional Influences

  • Long-term historical and regional processes can significantly influence species richness and diversity and will lead to exceptional patterns of diversity.
  • The Cape region of South Africa is smaller than the California Floristic Province but has more species.
  • Bond and Goldblatt attributed the Cape Floristic Region's unusually high species richness to several historic and geographic factors, including continental drift, a wide variety of soil types, repeated expansion, contraction, and isolation of plant populations, and refuge areas that reduced extinction rates.
  • Latham and Ricklefs (1993) studied temperate zone trees and found Europe, Eastern Asia, and Eastern North America all have roughly equal areas but vastly different levels of biological diversity.
  • Ralph (1985) found a negative correlation between foliage height diversity and bird species diversity in Argentina.
  • In Europe, mountains run east to west creating a barrier and cut-off for where trees could retreat.
  • Lower species richness is a consequence of a higher extinction rate.
  • The Appalachian Mountains in North America run north-south, giving temperate trees an avenue of retreat as temperatures become colder.
  • There are no mountain barriers in Asia.
  • Most temperate tree taxa originated in Eastern Asia before dispersing to Europe and North America.
  • After dispersal routes were cut, speciation continued in Asia.

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Explore geographic ecology, focusing on island size and species richness correlation. Understand MacArthur's definition, key factors influencing island biodiversity, and broad geographic patterns. Discover how historical events like Pleistocene climate warming shaped habitat distribution and species diversity.

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