Botany Chapter: Stems and Twigs
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What is the primary function of stems in plants?

Support and transport nutrients and water.

What type of growth do some plants exhibit?

  • Determinative growth
  • Yearly growth
  • Limited growth
  • Indeterminate growth (correct)
  • What are the two types of cambium mentioned?

    Vascular cambium and cork cambium.

    The area of the stem where leaves are attached is called a ______.

    <p>node</p> Signup and view all the answers

    Heartwood can conduct materials.

    <p>False</p> Signup and view all the answers

    Which type of stem stores food?

    <p>Tubers</p> Signup and view all the answers

    What do cotyledons provide for seedlings?

    <p>They store food needed by young seedlings.</p> Signup and view all the answers

    What are annual rings comprised of?

    <p>Spring and summer wood</p> Signup and view all the answers

    What is the role of vascular rays in a tree?

    <p>They function in lateral conduction of nutrients and water.</p> Signup and view all the answers

    What is a characteristic of hardwood?

    <p>Contains fibers and vessels</p> Signup and view all the answers

    Softwood comes from hardwood trees.

    <p>False</p> Signup and view all the answers

    What is the function of cork cambium?

    <p>To produce cork cells that reduce water loss.</p> Signup and view all the answers

    What is the term for the area of the stem where leaves are attached?

    <p>Node</p> Signup and view all the answers

    Which of the following describes a terminal bud?

    <p>Found at the tip of a twig</p> Signup and view all the answers

    What is the primary growth characteristic of stems with indeterminate growth?

    <p>They grow indefinitely.</p> Signup and view all the answers

    Which structure protects the buds on a woody twig?

    <p>Bud scales</p> Signup and view all the answers

    What term describes the angle between the petiole and the stem?

    <p>Axil</p> Signup and view all the answers

    Which of the following stem structures is distinguished by its ability to grow horizontally underground?

    <p>Rhizome</p> Signup and view all the answers

    What type of leaves are stipules commonly associated with?

    <p>Opposite leaves</p> Signup and view all the answers

    What do groups of bud scale scars on a twig indicate?

    <p>Age of the twig</p> Signup and view all the answers

    What is the main difference between monocotyledons and dicotyledons?

    <p>Monocots are typically annuals, while dicots can be annuals or perennials.</p> Signup and view all the answers

    Which type of stele features a solid core with phloem surrounding the xylem?

    <p>Protostele</p> Signup and view all the answers

    What is the primary function of lenticels in plant stems?

    <p>To facilitate gas exchange</p> Signup and view all the answers

    What types of plants primarily feature siphonosteles?

    <p>Ferns</p> Signup and view all the answers

    How does the vascular cambium function in herbaceous dicots?

    <p>It adds secondary xylem and secondary phloem.</p> Signup and view all the answers

    What occurs during the spring in terms of wood production in trees?

    <p>Larger vessel elements of secondary xylem are produced.</p> Signup and view all the answers

    What characterizes woody dicotyledonous stems compared to herbaceous stems?

    <p>They are primarily composed of secondary xylem.</p> Signup and view all the answers

    Which statement about annuals is correct?

    <p>They die after going from seed to maturity in one growing season.</p> Signup and view all the answers

    What is the role of the apical meristem in plant growth?

    <p>Increases stem length</p> Signup and view all the answers

    Which primary meristem is responsible for producing the epidermis?

    <p>Protoderm</p> Signup and view all the answers

    Which structure marks the food and water conducting tissue within leaf scars?

    <p>Bundle scars</p> Signup and view all the answers

    What do leaf and bud primordia develop into?

    <p>Mature leaves and buds</p> Signup and view all the answers

    What does the vascular cambium produce toward the surface of the plant?

    <p>Secondary phloem</p> Signup and view all the answers

    What type of cells does cork cambium primarily produce?

    <p>Cork cells with suberin</p> Signup and view all the answers

    Which part of the plant is protected by bud scales?

    <p>Apical meristem</p> Signup and view all the answers

    What type of cells are produced by the ground meristem?

    <p>Pith and cortex</p> Signup and view all the answers

    What primarily characterizes spring wood in conifers?

    <p>Larger tracheids than later in the season</p> Signup and view all the answers

    How do annual rings of wood indicate a tree's age?

    <p>Rings correspond to annual xylem production</p> Signup and view all the answers

    What is heartwood characterized by?

    <p>It consists of older, darker wood at the center</p> Signup and view all the answers

    What is the difference between sapwood and heartwood?

    <p>Sapwood contains living cells; heartwood does not</p> Signup and view all the answers

    What impact does environment have on the formation of annual rings in trees?

    <p>It influences the size of the rings produced</p> Signup and view all the answers

    What is primarily produced by the vascular cambium?

    <p>More secondary xylem than phloem</p> Signup and view all the answers

    Which element is absent in conifer wood?

    <p>Vessels</p> Signup and view all the answers

    What function do vascular rays serve in trees?

    <p>They facilitate lateral transport of nutrients and water</p> Signup and view all the answers

    What is the primary characteristic of hardwood compared to softwood?

    <p>It contains fibers and vessel elements.</p> Signup and view all the answers

    Which statement about heartwood is true?

    <p>A tree may still function after heartwood is removed.</p> Signup and view all the answers

    What distinguishes the vascular bundles of monocot stems from those of dicot stems?

    <p>Monocot stems contain no secondary vascular tissues.</p> Signup and view all the answers

    Which feature describes the bark of a tree?

    <p>It consists of alternating layers of crushed phloem and cork.</p> Signup and view all the answers

    What are laticifers in plants primarily involved in?

    <p>Secreting latex.</p> Signup and view all the answers

    Which statement accurately describes the structure of typical monocot vascular bundles?

    <p>They include two large vessels and several small vessels.</p> Signup and view all the answers

    Which of the following types of trees does not produce resin canals?

    <p>Monocot trees.</p> Signup and view all the answers

    What happens to xylem cells in monocots during their development?

    <p>They stretch and collapse, leaving air spaces.</p> Signup and view all the answers

    Study Notes

    Introduction to Stems

    • Most wood in household items comes from stems.
    • Grafting involves uniting parts of stems from different plants.
    • Some plants can grow indefinitely.

    External Form of A Woody Twig

    • Stems produce branches and leaves.
    • A woody twig has an axis with attached leaves.
    • Nodes are where leaves attach to the stem.
      • Alternate or spiral arrangement.
      • Opposite arrangement.
      • Whorled arrangement.
    • Internodes are the regions between nodes.
    • Leaves have flattened blades and are attached by petioles.

    Anatomy of A Woody Twig

    • Axil is the angle between petiole and stem.
    • Axillary buds are located in the axil.
      • Will develop into branches or flowers.
      • Bud scales protect buds.
    • Terminal buds are at the twig tip.
    • The growth makes the twig longer.
    • Bud scale scars indicate the age of the twig.
    • Stipules are leaf-like structures at the base of a leaf.

    Deciduous Tree and Shrubs

    • Deciduous trees lose all leaves annually.
    • Dormant axillary buds have leaf scars below.
    • Bundle scars mark food and water conducting tissue within leaf scars.

    Origin and Development of Stems

    • Apical meristem at the stem tip increases stem length.
    • It is dormant before the growing season begins.
    • Protected by bud scales and leaf primordia.
    • Leaf primordia are embryonic leaves.

    Primary Meristems

    • Apical meristem cells form three primary meristems:
      • Protoderm: Gives rise to the epidermis.
      • Procambium: Produces primary xylem and phloem.
      • Ground Meristem: Produces pith and cortex.

    Leaf Primordia and Bud Primordia

    • Leaf primordia and bud primordia develop into mature structures.
    • Traces branch off from xylem and phloem and enter leaves or buds.

    Vascular Cambium

    • Arises between primary xylem and primary phloem.
    • Produces secondary xylem towards the center and secondary phloem toward the surface.

    Cork Cambium

    • Produces cork cells with suberin and phelloderm cells.
    • Functions to reduce water loss and protect against injury.
    • Lenticels are parenchyma cells in cork for gas exchange.

    Tissue Patterns in Stems: Steles

    • Stele is the central cylinder of primary xylem, phloem, and pith.
    • Protostele: Solid core with phloem surrounding xylem.
      • Common in primitive plants.
    • Siphonosteles: Tubular with pith in the center.
      • Typical in ferns.
    • Eusteles: Discrete vascular bundles.
      • Found in flowering plants and conifers.

    Seed Leaves

    • Cotyledons are seed leaves.
    • Dicotyledons (dicots) have two cotyledons.
    • Monocotyledons (monocots) have a single cotyledon.

    Herbaceous Dicotyledonous Stems

    • Annuals die after completing their life cycle within one season.
    • Most monocots and many dicots are annuals.

    Tissue Patterns in Herbaceous Dicotyledonous Stems

    • Herbaceous dicots have vascular bundles arranged in a cylinder.
    • Vascular cambium adds secondary xylem and phloem.

    Woody Dicotyledonous Stems

    • Secondary xylem is wood.
    • Vascular and cork cambium active all year:
      • Produces ungrained, uniform wood.
    • Some tropical trees produce uniform wood.

    Seasonal Production of Wood

    • Spring wood has large vessel elements.
    • Summer wood has fewer, smaller vessel elements.
    • Alternating spring and summer wood creates rings.
    • Annual ring reflects the xylem growth in one year.

    Conifers

    • Conifers lack vessels and fibers.
    • Tracheids are larger in the spring.

    Annual Rings and Vascular Rays

    • One year of xylem growth forms an annual ring.
    • Annual rings indicate the tree's age and climate changes.
    • Vascular rays are parenchyma cells for lateral nutrient and water transport.
      • Xylem ray is within the xylem.
      • Phloem ray is in the phloem.

    Cross Section of a Young Stem

    • Diagram highlighting secondary growth in a young stem.

    A Block of a Woody Dicotyledonous Stem

    • 3D view of dicot wood.

    Heartwood and Sapwood

    • Heartwood is the older, darker, non-functioning wood in the center.
    • Sapwood is the lighter, active wood in the outer layers.

    Removal of Wood

    • Heartwood supports the tree but doesn't conduct materials.
    • It is possible to remove portions of sapwood without destroying the tree.

    Softwood and Hardwood

    • Softwood is wood from conifers.
    • Hardwood is wood from dicot trees.
    • Resin canals are tubes in the wood, typically present in softwoods.

    Bark and Laticifers

    • Bark consists of tissues outside the vascular cambium, including secondary phloem.
    • Mature bark alternates between crushed phloem and cork layers.
    • Laticifers are ducts in phloem that contain latex.
      • Latex can be used to produce rubber, chicle, or morphine.

    Monocotyledonous Stems

    • Monocots lack vascular and cork cambium.
    • They do not produce secondary vascular tissues or cork.
    • Vascular bundles are scattered throughout the stem.

    Typical Monocot Vascular Bundle

    • Contains large and small vessels.
    • Phloem consists of sieve tubes and companion cells.
    • Vascular bundles are surrounded by sclerenchyma cells.

    Specialized Stems

    • Rhizomes are horizontal underground stems.
    • Runners are horizontal stems that grow above ground.
    • Stolons are produced beneath the ground and grow in different directions.

    Tubers and Bulbs

    • Tubers are swollen underground stems that store food.
    • Bulbs are buds surrounded by fleshy storage leaves with a small stem at the base.

    Corms and Cladophylls

    • Corms resemble bulbs but are mostly stem tissue.
    • Cladophylls are flattened, leaf-like stems.

    Wood and Its Uses

    • Half of the wood's weight is water content.
    • The dry part is mainly composed of cellulose and lignin.
    • Density is the weight per unit volume.
    • Durability is the resistance to decay.

    Types of Sawing

    • Boards cut radially show annual rings in side view.
    • Boards cut tangentially show annual rings as irregular bands.

    Knots

    • Knots are the bases of lost branches covered by annual rings.
    • They are more common in older parts of a log.

    Wood Products

    • Lumber is used primarily for construction.
    • Sawdust and waste is used for particleboard and pulp.
    • Veneer is a thin layer of wood glued to cheaper lumber.
    • Pulp is used for paper, synthetic fibers, and plastics.
    • Wood is also used as fuel in developing countries.

    Introduction to Stems

    • Stems provide the structural support for plants and are the source of most wood used in daily life.
    • Grafting, a technique involving the joining of stems from different plant varieties, has been practiced for centuries.
    • Some plants possess indeterminate growth, allowing them to continue growing indefinitely.

    External Form of a Woody Twig

    • Stems develop a shoot system with branches and leaves.
    • A woody twig consists of an axis with attached leaves.
    • Nodes represent areas where leaves attach to the stem, exhibiting various arrangements: alternate, opposite, or whorled.
    • Internodes are the sections of stem between nodes.
    • Leaves typically have a flattened blade and are attached to the twig via a petiole.

    Anatomy of a Woody Twig

    • The axil is the angle formed between the petiole and the stem where an axillary bud is located.
    • Axillary buds develop into branches or flowers in flowering plants and are protected by bud scales.
    • The terminal bud is situated at the tip of the twig and contributes to stem elongation.
    • Stem age can be determined by counting groups of bud scale scars.
    • Stipules are paired structures, often leaf-like, located at the base of a leaf.

    Deciduous Trees and Shrubs

    • Deciduous plants shed all their leaves annually.
    • After leaf fall, dormant axillary buds remain with leaf scars below.
    • Bundle scars mark the location of food and water conducting tissue within leaf scars.

    Origin and Development of Stems

    • The apical meristem, located at the stem tip, contributes to stem length and is protected by bud scales and leaf primordia.
    • Leaf primordia are tiny embryonic leaves that develop into mature leaves.

    Primary Meristems

    • Apical meristem cells differentiate into three primary meristems: protoderm, procambium, and ground meristem.
    • Protoderm gives rise to the epidermis.
    • Procambium produces primary xylem and phloem.
    • Ground meristem forms pith and cortex, both composed of parenchyma cells.

    Leaf Primordia and Bud Primordia

    • Leaf primordia and bud primordia develop into mature leaves and buds, respectively.
    • Traces, strands of xylem and phloem, branch off from the vascular cylinder and extend into leaves or buds.

    Vascular Cambium

    • The vascular cambium, a meristematic band, arises between primary xylem and primary phloem.
    • The vascular cambium produces secondary xylem towards the center and secondary phloem towards the surface.

    Cork Cambium

    • Cork cambium (phellogen) produces cork cells containing suberin and phelloderm cells.
    • Cork cells contribute to water loss reduction and stem protection against injury.
    • Lenticels are parenchyma cells within cork that facilitate gas exchange.

    Tissue Patterns in Stems: Steles

    • The stele, the central cylinder of the stem, comprises primary xylem, primary phloem, and pith (if present).
    • Protostele, a solid core with phloem surrounding xylem, is found in primitive seed plants, whisk ferns, club mosses, and ferns.
    • Siphonosteles have a tubular structure with pith in the center, common in ferns.
    • Eusteles, with discrete vascular bundles, are characteristic of flowering plants and conifers.

    Seed Leaves

    • Cotyledons are seed leaves attached to the embryonic stem, serving as food storage for the developing seedling.
    • Dicotyledons (dicots) are flowering plants with two cotyledons in their seeds.
    • Monocotyledons (monocots) are flowering plants with a single cotyledon in their seeds.

    Herbaceous Dicotyledonous Stems

    • Annuals are plants that complete their life cycle from seed to maturity within a single growing season.
    • They are typically green and herbaceous and rely primarily on primary tissues.
    • Though many dicots are annuals, most monocots belong to this category.

    Tissue Patterns in Herbaceous Dicotyledonous Stems

    • Herbaceous dicots possess discrete vascular bundles arranged in a cylinder.
    • The vascular cambium arises between primary xylem and primary phloem, generating secondary xylem and phloem growth.

    Woody Dicotyledonous Stems

    • Wood is synonymous with secondary xylem.
    • Variations in wood development are influenced by the continuous activity of vascular cambium and cork cambium throughout the year.
    • Tropical trees often produce ungrained, uniform wood due to consistent growth.

    Seasonal Production of Wood

    • In spring, the vascular cambium produces relatively large vessel elements of secondary xylem, forming spring wood.
    • Later in the season, fewer and smaller vessel elements are produced, leading to the formation of summer wood.
    • The alternating pattern of spring and summer wood creates light and dark rings, known as annual rings.
    • Environmental factors impact the size of these rings.
    • Xylem growth within a year constitutes an annual ring.

    Conifers

    • Conifers lack vessels and fibers, but their tracheids are larger in spring than later in the season.

    Annual Rings and Vascular Rays

    • An annual ring represents a year's worth of xylem growth.
    • The vascular cambium produces more secondary xylem than phloem, forming the bulk of the tree trunk.
    • Annual rings provide insights into a tree's age and the climate during its lifetime.
    • Vascular rays, composed of parenchyma cells, function in lateral conduction of water and nutrients.
    • Vascular rays extend through both xylem and phloem, forming xylem rays and phloem rays, respectively.

    Cross Section of a Young Stem

    • A cross section of a young stem with secondary growth reveals distinct tissue layers.

    A Block of a Woody Dicotyledonous Stem

    • A 3-D view of dicot wood showcases the arrangement of vascular bundles, annual rings, and vascular rays.

    Heartwood and Sapwood

    • Heartwood, the older, darker wood at the center of the stem, accumulates resins, gums, and tannins.
    • Sapwood, the lighter-colored, functional xylem closest to the cambium, actively conducts water and nutrients.

    Removal of Wood

    • Heartwood primarily provides structural support and is no longer involved in material transport.
    • Removal of heartwood does not affect the tree's functionality.
    • Portions of sapwood can also be removed without harming the tree.

    Softwood and Hardwood

    • Softwood refers to the wood of conifers, lacking fibers or vessel elements.
    • Hardwood, derived from dicot trees, contains fibers and vessel elements.
    • Resin canals, tube-like structures scattered throughout xylem and other tissues, are typically found in conifers.

    Bark and Laticifers

    • Bark denotes tissues external to the vascular cambium, including secondary phloem.
    • Mature bark consists of alternating layers of crushed phloem and cork.
    • Laticifers, ducts primarily located in phloem, contain latex-secreting cells which are used in the production of rubber, chicle, and morphine.

    Monocotyledonous Stems

    • Monocot stems lack a vascular cambium and a cork cambium, resulting in the absence of secondary vascular tissues or cork.
    • Primary xylem and phloem are organized in scattered vascular bundles throughout the stem.

    Typical Monocot Vascular Bundle

    • Monocot vascular bundles typically contain two large vessels and several small vessels.
    • Early formed xylem cells elongate and collapse, leaving irregularly shaped air spaces.
    • Phloem comprises sieve tubes and companion cells.
    • The vascular bundle is surrounded by a sheath of sclerenchyma cells.

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